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Anolis marmoratus DUMÉRIL & BIBRON, 1837

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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
SubspeciesAnolis marmoratus alliaceus COPE 1864
Anolis marmoratus caryae LAZELL 1964
Anolis marmoratus girafus LAZELL 1964
Anolis marmoratus inornatus LAZELL 1964
Anolis marmoratus marmoratus DUMÉRIL & BIBRON 1837
Anolis marmoratus setosus LAZELL 1964
Anolis marmoratus speciosus (GARMAN 1887) 
Common NamesE: Guadeloupe Anole, Guadeloupean anole, Leopard Anole 
SynonymAnolis marmoratus DUMÉRIL & BIBRON 1837: 139
Anolis speciosus GARMAN 1888
Anolis marmoratus — UNDERWOOD in WILLIAMS et al. 1959
Ctenonotus marmoratus — SCHWARTZ & HENDERSON 1988: 113
Anolis marmoratus — SCHWARTZ & HENDERSON 1991: 297
Ctenonotus marmoratus — NICHOLSON et al. 2012
Ctenonotus marmoratus — NICHOLSON et al. 2018

Anolis marmoratus alliaceus (COPE 1864)
Anolis alliaceus COPE 1864: 175
Anolis alliaceus — GÜNTHER 1888: 363
Anolis bimaculatus alliaceus — UNDERWOOD in WILLIAMS et al. 1959
Anolis marmoratus alliaceus — LAZELL 1964: 374
Anolis marmoratus alliaceus — SCHMIDT 1981
Ctenonotus marmoratus alliaceus — SCHWARTZ & HENDERSON 1988: 114
Anolis marmoratus alliaceus — HENKEL 2006
Ctenonotus marmoratus alliaceus — NICHOLSON et al. 2012
Ctenonotus marmoratus alliaceus — NICHOLSON et al. 2018

Anolis marmoratus caryae LAZELL 1964
Ctenonotus marmoratus caryae — SCHWARTZ & HENDERSON 1988
Anolis caryae — ROUGHGARDEN 1995
Anolis terraealtae caryae — POWELL et al. 2012
Ctenonotus marmoratus caryae — NICHOLSON et al. 2012
Ctenonotus marmoratus caryae — NICHOLSON et al. 2018
Anolis marmoratus caryae — HEDGES et al. 2019

Anolis marmoratus girafus LAZELL 1964
Anolis marmoratus girafus — SCHMIDT 1981
Ctenonotus marmoratus girafus — SCHWARTZ & HENDERSON 1988
Ctenonotus marmoratus girafus — NICHOLSON et al. 2012
Ctenonotus marmoratus girafus — NICHOLSON et al. 2018

Anolis marmoratus inornatus LAZELL 1964
Anolis marmoratus inornatus — SCHMIDT 1981
Ctenonotus marmoratus inornatus — SCHWARTZ & HENDERSON 1988
Ctenonotus marmoratus inornatus — NICHOLSON et al. 2012
Ctenonotus marmoratus inornatus — NICHOLSON et al. 2018

Anolis marmoratus setosus LAZELL 1964
Anolis marmoratus setosus — SCHMIDT 1981
Ctenonotus marmoratus setosus — SCHWARTZ & HENDERSON 1988
Ctenonotus marmoratus setosus — NICHOLSON et al. 2012
Ctenonotus marmoratus setosus — NICHOLSON et al. 2018

Anolis marmoratus speciosus (GARMAN 1887)
Anolis speciosus GARMAN 1887: 45
Anolis speciosus speciosus — UNDERWOOD in WILLIAMS et al. 1959
Anolis marmoratus speciosus — LAZELL 1964: 384
Ctenonotus marmoratus speciosus — SCHWARTZ & HENDERSON 1988: 115
Anolis marmoratus speciosus — HESELHAUS & SCHMIDT 1995
Anolis marmoratus speciosus — BREUIL et al. 2009
Ctenonotus marmoratus speciosus — NICHOLSON et al. 2012
Ctenonotus marmoratus speciosus — NICHOLSON et al. 2018 
DistributionLesser Antilles: Guadeloupe, Ile Desecheo Saintes, La Desirade,
French Guiana (introduced)

marmoratus: Guadeloupe: plain of Capesterre in southeastern Basse-Terre; intergrades with A. m. alliaceus, A. m. girafus, and A. m. speciosus.

alliaceus: central highlands in the Basse-Terre portion of Guadeloupe; intergrades with A. m. girafus, A. m. setosus, and A. m. speciosus.

caryae: Terre-de-Bas, Îles des Saintes; intergrades with A. m. terraealtae; introduced at Cayenne, Guiane Française.

girafus: leeward coast of Basse-Terre portion of Guadeloupe, from vicinity of Malendure and adjacent Ilets de Pigeon, south to Ballif; intergrades with A. m. setosus.

inornatus: northern Grande-Terre; Ilet Macou; intergrades with A. m. speciosus.

setosus northern coast of Basse-Terre portion of Guadeloupe from vicinity of Deshaies to Ste.-Rose; intergrades with A. m. speciosus.

speciosus: southwestern portion of Grande-Terre portion of Guadeloupe, onto isthmus connecting Grande-Terre and Basse-Terre; Ilet du Cochons, Ilet du Gosier, Ilet Christophe. Introduced to French Guiana (HOOGMOED, pers. comm.).

Type locality: Martinique (in error); revised to Capesterre on the Basse-Terreportion of Guadeloupe by Lazell, 1964.  
Reproductionoviparous 
TypesSyntypes: MNHN-RA 0794 (lectotype), MNHN-RA 5491 (paralectotype)
Holotype: BMNH 1946.8.28.96 [alliaceus]
Holotype: MCZ 70666 [caryae]
Holotype: MCZ 71259 [girafus]
Holotype: MCZ 71036 [inornatus]
Holotype: MCZ 70813 [setosus]
Lectotype: MCZ 6172 [speciosus] 
Diagnosis 
CommentSubspecies: Anolis marmoratus ferreus, terraealtae, chrysops, desiradei, and kahouannensis are now considered a valid species.

Species group: Ctenonotus bimaculatus species group (fide NICHOLSON et al. 2012).

Hybridization: Male A. marmoratus and female A. ferreus can interbreed and produce offspring (Weidlich 2023, Anole Annals) 
EtymologyNamed after Latin marmoratus, a, um, from the participle of marmoro, ‘covered with marble, marbled’, in reference to the marbling on the head and neck.

A. m. alliaceus was named after Latin allium or alium, ‘garlic’and the suffix -aceus, a, um, ‘which resembles to, which has the nature of’. Cope (1864) did not explain his choice (Fretey 2023).

A. m. caryae was named after Margaret M. Cary who sponsored the first trip of Lazell to the Caribbean (Fretey 2023).

A. m. girafus has no known etymology and may be an arbitrary combination of letters. Lazell (1964: 337) indicates that “The name girafus has been invented
to indicate the reticulate pattern” (Fretey 2023).

A. m. inornatus was named after Latin inornatus, a, um, = ‘unadorned (Fretey 2023).

A. m. setosus was named after Latin setosus (or saetosus), a, um, ‘bristly, covered with hair’ (Fretey 2023).

A. m. speciosus was named after Latin speciosus, a, um, = ‘of beautiful appearance’ (Fretey 2023). 
References
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