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Anolis meridionalis BOETTGER, 1885

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards)
Common NamesPortuguese: Calango-Bandeira, Lagarto-Preguiça, Papa-Vento 
SynonymAnolis meridionalis BOETTGER 1885: 437
Anolis holotropis BOULENGER 1895: 522
Norops sladeniae BOULENGER 1903: 69
Anolis steinbachi GRIFFIN 1917 (fide VANZOLINI & WILLIAMS 1970)
Norops marmorata AMARAL 1932:63
Anolis chrysolepis meridionalis — HELLMICH 1960:22 (partim)
Anolis meridionalis — PETERS et al. 1970: 60
Anolis meridionalis — DIRKSEN & DE LA RIVA 1999
Norops meridionalis — NICHOLSON 2002
Anolis meridionalis — D’ANGIOLELLA et al. 2011
Norops meridionalis — NICHOLSON et al. 2012
Norops meridionalis — CACCIALI et al. 2016
Norops meridionalis — NICHOLSON et al. 2018 
DistributionBrazil (Mato Grosso, Goias, São Paulo), Paraguay, Bolivia (Beni, Santa Cruz)

Type locality: Paraguay. Neotype locality: Colonia Ybycui, Estancia Ybycui, Departamento Canindeyú, Paraguay, (designation by Motte and Cacciali, 2009)  
TypesNeotype: MNHNP (Paraguay), desgnated by MOTTE & CACCIALI 2009
Holotype: lost
Holotype: unlocated fide D’Angiolella et al. 2011 [holotropis]
Holotype: unlocated fide D’Angiolella et al. 2011 [sladeniae]
Holotype: CM 988 [steinbachi]
Holotype: MZUSP 737 [marmorata] 
DiagnosisDescription. Maximum SVL 56 mm. Specimens analyzed presented vertebral scales anteriorly small, keeled, posteriorly increasing in size and forming a gradual transition toward the granules on flanks; they do not form rows of enlarged scales along dorsum. The neotype description provided by Motte and Cacciali (2009) states that vertebral scales form eight rows of enlarged scales along back that are distinctly larger than scales on the flanks. Scales on upper arm keeled, markedly larger than vertebral scales. Scales on snout anteriorly weakly to distinctly keeled, uni- or multi-carinate and slightly larger than the flat posterior scales. Supraorbital semicircles with enlarged, weakly keeled scales. Supraocular region with a group of distinctly enlarged scales surrounded posteriorly and laterally by granules and anteriorly by smaller scales. Interparietal scale distinctly larger than adjacent scales. Loreals 6–7; canthals 5–6; 8–10 scales between second canthals; 0– 1 scales between supraorbital semicircles; 0– 2 scales between supraorbital semicircles and interparietal, postmentals 5–8 [from D’ANGIOLELLA et al. 2011]. 
CommentSynonymy after Peters et al. 1970, VANZOLINI & WILLIAMS 1970, and D’ANGIOLELLA et al. 2011.

N. meridionalis may be composed by at least five cryptic species (Guarnizo et al. 2015). However, they do not make any formal changes.

Species group: Norops auratus Species Group (fide Nicholson et al. 2012) 
  • Araujo, Cybele O.; Fábio Maffei, Décio T. Corrêa, Guilherme M. Moya, Bruno T. M. Nascimento, Selma Maria A. Santos 2014. Lizards from Estação Ecológica de Santa Bárbara, a remnant of Cerrado in the state of São Paulo, Brazil. Check List 10 (5): 1038-10 - get paper here
  • Barbour, T. 1934. The anoles. II. The mainland species from México southward. Bull. Mus. Comp. Zool. Harvard 77: 121-155. - get paper here
  • Boettger, O. 1885. Berichtigung der Liste von Reptilien und Batrachiern aus Paraguay. Zeitschr. Naturw. 58: 436-437 - get paper here
  • Boulenger, George A. 1895. Description of a new Anolis from Brazil. Ann. Mag. nat. Hist. (6)15 (90): 522-523 - get paper here
  • Boulenger, George A. 1903. List of the batrachians and reptiles collected by M. A. Robert at Chapadá, Matto Grosso, and presented by Mrs. Percy Sladen to the British Museum. (Percy Sladen Expedition to Central Brazil). Proceedings of the Zoological Society of London 1903, 2 (1): 69-70 - get paper here
  • Bressan, Paulo Magalhães; Maria Cecília Martins Kierulff, Angélica Midori Sugieda (eds.) 2009. FAUNA AMEAÇADA DE EXTINÇÃO NO ESTADO DE SÃO PAULO: VERTEBRADOS. São Paulo: Fundação Parque Zoológico de São Paulo: Secretaria do Meio Ambiente, 2009. - get paper here
  • Cacciali, Pier; Norman J. Scott, Aida Luz Aquino Ortíz, Lee A. Fitzgerald, and Paul Smith 2016. The Reptiles of Paraguay: Literature, Distribution, and an Annotated Taxonomic Checklist. SPECIAL PUBLICATION OF THE MUSEUM OF SOUTHWESTERN BIOLOGY, NUMBER 11: 1–373 - get paper here
  • D'angiolella, Annelise B.; Tony Gamble, Teresa C. S. Avila-Pires, Guarino R. Colli, Brice P. Noonan, and Laurie J. Vitt 2011. Anolis chrysolepis Duméril and Bibron, 1837 (Squamata: Iguanidae), Revisited: Molecular Phylogeny and Taxonomy of the Anolis chrysolepis Species Group. Bull. Mus. Comp. Zool. Harvard 160 (2): 35-63. - get paper here
  • Dirksen, L. & De la Riva, I. 1999. The lizards and amphisbaenians of Bolivia (Reptilia, Squamata): checklist, localities, and bibliography. Graellsia 55: 199-215 - get paper here
  • Gonzalez R. C. et al. 2020. Lista dos Nomes Populares dos Répteis no Brasil – Primeira Versão. Herpetologia Brasileira 9 (2): 121 – 214 - get paper here
  • Griffin, L. E. 1917. A list of the South American lizards of the Carnegie Museum, with descriptions of four new species. Annals of the Carnegie Museum 11: 304-320. - get paper here
  • Guarnizo, Carlos E.; Fernanda P. Werneck, Lilian G. Giugliano, Marcella G. Santos, Jéssica Fenker, Lucas Sousa, Annelise B. D’Angiolella, Adriana R. dos Santos, Christine Strüssmann, Miguel T. Rodrigues, Tainá F. Dorado-Rodrigues, Tony Gamble, Guarin 2015. Cryptic lineages and diversification of an endemic anole lizard (Squamata, Dactyloidae) of the Cerrado hotspot. Molecular Phylogenetics and Evolution 94: 279-289, doi:10.1016/j.ympev.2015.09.005 - get paper here
  • Guerra, V., Ramalho, W. P., Machado, I. F., & Brandão, R. A. 2022. Herpetofauna of the Serra do Tombador Nature Reserve, State of Goiás, Central Brazil. Arquivos de Zoologia, 53(3), 33-51 - get paper here
  • MARQUES, O. A. V. et al. 2009. Repteis. In: Bressan et al., FAUNA AMEAÇADA DE EXTINÇÃO NO ESTADO DE SÃO PAULO: VERTEBRADOS. São Paulo: Fundação Parque Zoológico de São Paulo: Secretaria do Meio Ambiente, pp. 285-327 - get paper here
  • Motte, Martha; Pier Cacciali 2009. Descripción de un neotipo para Anolis meridionalis Boettger, 1885 (Sauria: Polychrotidae). Cuadernos de Herpetología 23 (1):19-24 - get paper here
  • Nicholson KE, C. Guyer, and JG Phillips 2017. Biogeographic Origin of Mainland Norops (Squamata: Dactyloidae). Assumptions Inhibiting Progress in Comparative Biology (eds. Crother and Parenti), pp. 169–184
  • Nicholson, K.E. 2002. Phylogenetic analysis and a test of the current infrageneric classification of Norops (beta Anolis). Herpetological Monographs 16: 93-120 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Nogueira, Cristiano; Guarino R. Colli, Gabriel Costa & Ricardo B. Machado 2010. Diversidade de répteis Squamata e evolução do conhecimento faunístico no Cerrado. - get paper here
  • Peters, James A. & Donoso-Barros, Roberto 1970. Catalogue of the Neotropical Squamata: Part II. Lizards and Amphisbaenians. Bull. US Natl. Mus. 297: 293 pp. - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Ramalho, Werther Pereira; Jefferson Rocha Silva, Philip Teles Soares, Denes Ferraz, Filipe Viegas Arruda, Vitor Hugo Mendonça do Prado 2018. The anurans and squamates of a peri-urban Cerrado remnant in the State of Goiás, Central Brazil. Herpetology Notes 11: 573-583 - get paper here
  • Ramalho, Werther Pereira; Vinícius Guerra Batista; Luciano Roberto Passos Lozi 2014. Anfíbios e répteis do médio rio Aporé, estados de Mato Grosso do Sul e Goiás, Brasil. Neotropical Biology and Conservation 9 (3): 147-160; doi: 10.4013/nbc.2014.93.04 - get paper here
  • RIBEIRO-JÚNIOR, MARCO A. 2015. Catalogue of distribution of lizards (Reptilia: Squamata) from the Brazilian Amazonia. I. Dactyloidae, Hoplocercidae, Iguanidae, Leiosauridae, Polychrotidae, Tropiduridae. Zootaxa 3983 (1): 001–110 - get paper here
  • Ribeiro-Júnior, Marco A. & Silvana Amaral 2016. Diversity, distribution, and conservation of lizards (Reptilia: Squamata) in the Brazilian Amazonia. Neotropical Biodiversity, 2:1, 195-421 - get paper here
  • Uetanabaro, Masao; Franco Leandro Souza, Paulo Landgref Filho,<br />Arlindo Figueira Beda & Reuber Albuquerque Brandão 2007. Anfíbios e répteis do Parque Nacional da Serra da Bodoquena, Mato Grosso do Sul, Brasil. Biota Neotropica 7 (3) - get paper here
  • Vanzolini,P.E. & Williams,E.E. 1970. South American anoles: the geographic differentiation and evolution of the Anolis chrysolepis species group (Sauria, Iguanidae). Arquivos de Zoologia (São Paulo) 19: 1-298 - get paper here
  • Vechio, Francisco dal; Renato Recoder; Miguel Trefaut Rodrigues; Hussam Zaher 2013. The herpetofauna of the estação ecológica de uruçuí-una, state of Piauí, Brazil. Pap. Avulsos Zool. 53(16): 225‐243 - get paper here
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