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Anolis quadriocellifer BARBOUR & RAMSDEN, 1919

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards)
Common NamesE: Cuban Eyespot Anole, Peninsula Anole 
SynonymAnolis quadriocellifer BARBOUR & RAMSDEN 1919:158
Anolis calliurus AHL 1924: 249
Anolis homolechis quadriocellifer — BARBOUR 1937: 127
Anolis quadriocellifer — SCHWARTZ & HENDERSON 1991: 321
Anolis quadriocellifer — RODRÍGUEZ SCHETTINO 1999: 341
Norops quadriocellifer — NICHOLSON 2002
Norops quadriocellifer — NICHOLSON et al. 2012
Norops quadriocellifer — NICHOLSON et al. 2018 
DistributionW Cuba (Guanahacabibes)

Type locality: Cabo de San Antonia, Ensenada de Cajón, Pinar del Rio Province, Cuba.  
TypesHolotype: MCZ 11867. 
DiagnosisDESCRIPTION: Size moderate (SVL in males to 55 mm, in females to 40 mm); head scales all sharply keeled; total loreals 16-42; scales between supraorbitals 0-2 (mode 1); 1-4 (mode 3) scales between interparietal and supraorbital semicircles; 6 postrostrals; 2-7 (mode 4) postmentals; suboculars in contact with supralabials; scales on posterior margin of interparietal large, sharply demarcated from dorsals; dorsals small, granular, 2 middorsal rows slightly enlarged; ventrals smooth (occasionally keeled), large, cycloid, strongly imbricate, in diagonal and transverse rows; supradigital scales usually multicarinate, very occasionally smooth; tail laterally compressed, crested, verticillate. 7 lateral scales/verticil, 5 middorsals/ verticil, on crest 3/verticil. Dorsum some shade of brown to tan; usually a pair of large. triangular, black scapular patches, outlined with white, and at times with a single, white, central dot, but patch at times not present or reduced and brown; occasionally a second pair of black ocelli present on anterior body; 3-4 vertical, white flank bars composed of white dots; 1-2 white or white-dashed flank lines, the upper at level of bottom of triangular scapular patch; dewlap variable, yellow, yellow-orange, orange, in some populations with 3 red bars; females similar to males but with about 4 pale dorsal rhombs, the scapular patches located in the first rhombal constriction (Schwartz & Henderson 1991: 321). 
CommentFor illustrations see Barbour and Ramsden, 1919; Ruibal and Williams, 1961; Schwartz, 1968.

Species group: Norops sagrei species group (fide NICHOLSON et al. 2012). 
  • Ahl, E. 1924. Neue Reptilien und Batrachier aus dem zoologischen Museum Berlin. Archiv für Naturgeschichte 90 (A) 5: 246-254
  • Barbour, T. and C. T. Ramsden. 1919. The herpetology of Cuba. Mem. Mus. Comp. Zool., Harvard Coll. 47: 69-213. - get paper here
  • Barbour,T. 1937. Third list of Antillean reptiles and amphibians. Bull. Mus. comp. Zool. Harvard 82 (2): 77-166 - get paper here
  • Fläschendräger, Axel 2017. Beobachtungen an einigen Anolis-Arten im westlichen Kuba. Terraria-Elaphe 2017 (5): 52-63 - get paper here
  • Muñoz, Martha M.; Johanna E. Wegener and Adam C. Algar 2014. Untangling Intra- and Interspecific Effects on Body Size Clines Reveals Divergent Processes Structuring Convergent Patterns in Anolis Lizards. The American Naturalist 184 (5), DOI: 10.1086/678084 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Rodríguez Schettino, Lourdes (ed.) 1999. The iguanid lizards of Cuba. University Press of Florida, 428 pp.
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • Ruibal, R. and E. E. Williams. 1961. The taxonomy of the Anolis homolechis complex of Cuba. Bull. Mus. Comp. Zool. Harvard 125: 211-246. - get paper here
  • Schwartz, A. 1968. The Cuban lizards of the Anolis homolechis complex. Tulane Studies in Zoology 14: 140-184. - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
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