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Anolis salvini BOULENGER, 1885

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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
Common NamesSalvin's Anole (vociferans: Koanng Anole) 
SynonymAnolis salvini BOULENGER 1885: 75
Anolis salvini — GÜNTHER 1885: 48
Anolis vociferans MYERS 1971: 10 (fide KÖHLER 2007)
Anolis pentaprion — WALTERS 1953 (not COPE)
Norops vociferans — VILLA et al. 1988
Norops vociferans — KÖHLER 2000: 64
Norops vociferans — NICHOLSON 2002
Norops vociferans — SAVAGE 2002: 478
Norops salvini — NICHOLSON 2002
Norops salvini — NICHOLSON et al. 2012
Norops salvini — NICHOLSON et al. 2018 
DistributionPanama, Costa Rica, Guatemala ?

Type locality: Guatemala (in error fide KÖHLER 2007).

vociferans: Costa Rica, Panama (pacific side of the Cordillera de Talamanca from Cerro de la Muerte to Volcan de Chiriqui).Type locality: Panama,Chiriqui, 4 km W Cerro Punta, 1829 m elevation.  
TypesHolotype: BMNH 1946.9.8.19 (formerly
Holotype: Holotype: AMNH 69621 [vociferans] 
DiagnosisDEFINITION AND DIAGNOSIS (vociferans): Medium-size (snout to vent length 57 mm. in largest specimen), short-legged anole of the beta group, having trans-verse processes on some autotomous caudal vertebrae. Brown or gray-brown above with variable, in some cases partly reticulate, dark pattern usually including interorbital bar, short postocular stripe, V-shaped marking between forelimbs, and butterfly-shaped mark on base of tail between hind limbs; red dewlap, bright blue sliver of color at corners of nmouth, and black throat lining. Trunk granulation variable: granules small and convex on sides, sometimes larger and flattened on middorsum, either without definite arrangement or weakly disposed in horizontal rows dorsally and in oblique rows laterally; some dorsal granules keeled; ventral granules ontogenetically changing from conical and juxtaposed to keeled and imbricate. Middorsal caudal scales carinate but not pro jected in dorsal crest. Plates atop head relatively large, frontal ridges virtually absent, canthal ridges distinct, elongated supraciliary scale(s) present. Male with pair of moderately enlarged postanal scales.
Anolis vociferans is most likely to be confused with the related A. pentaprion, but vociferans can be distinguished by the presence of a dark interorbital bar, enlarged postanal scales in males, keeling on scattered middorsal granules (and keeled ventrals in older specimens), and fewer supralabials to center of eye (vociferans, 6-8, usually 7 at least on one side; pentaprion, 7-10, usually 9 at least on one side). The species also differs from southern populations of pentaprion in that the middorsal caudal scales, although carinate, are not projected in a low, serrated crest. The absence of bony parietal projections, presence of enlarged postanal scales in males, different caudal scutellation, and a very different color pattern, among other characters, easily distinguishes Anolis vociferans from its smaller highland relative, A. fungosus (from MYERS 1971). 
CommentNot listed by KÖHLER 2000.

Species group: Norops auratus Species Group (fide Nicholson et al. 2012). Other authors have stated that A. vociferans belongs to the pentaprion group which includes seven species: fungosus, ibague, ortonii, pentaprion, sulcifrons, utilensis, and vociferans. 
EtymologyNamed after the Latin “vocifer” = “to cry aloud”, in allusion to the squeaking noises made by specimens when they are first captured. 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bienentreu, J.-F.; A. Hertz; G. Köhler; S. Lotzkat. 2013. Distribution extension for Anolis salvini Boulenger, 1885 (Reptilia: Squamata: Dactyloidae), in western Panama. Check List 9 (1):169-174 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Echelle,A.A.; Echelle,A.F. & FITCH,H.S. 1971. A new anole from Costa Rica. Herpetologica 27: 354-362 - get paper here
  • Günther, A. C. L. G. 1885. Reptilia and Batrachia. Biologia Centrali-Américana. Taylor, & Francis, London, 326 pp. [published in parts from 1885-1902; reprint by the SSAR 1987] - get paper here
  • Köhler G. 1996. A new species of anole of the Norops pentaprion group from Isla de Utila, Honduras. (Reptilia: Sauria: Iguanidae) [Norops utilensis]. Senckenbergiana biologica 75 (1/2): 23-31.
  • Köhler, G. 2000. Reptilien und Amphibien Mittelamerikas, Bd 1: Krokodile, Schildkröten, Echsen. Herpeton Verlag, Offenbach, 158 pp.
  • Köhler, G. 2008. Reptiles of Central America. 2nd Ed. Herpeton-Verlag, 400 pp.
  • Köhler, G. 2010. A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae). Zootaxa 2354: 1–18 - get paper here
  • Köhler, Gunther 2007. Assessing the status of Anolis salvini Boulenger 1885 and A. bouvierii Bocourt 1873 based on the primary types (Reptilia, Squamata, Polychrotidae). Senckenbergiana Biologica 87(1):125-130 (1-6)
  • KÖHLER, GUNTHER 2014. Characters of external morphology used in Anolis taxonomy—Definition of terms, advice on usage, and illustrated examples. Zootaxa 3774 (2): 201–257 - get paper here
  • Köhler, Gunther; Marcos Ponce, Javier Sunyer, and Abel Batista 2007. FOUR NEW SPECIES OF ANOLES (GENUS ANOLIS) FROM THE SERRANÍA DE TABASARÁ, WEST-CENTRAL PANAMA (SQUAMATA: POLYCHROTIDAE). Herpetologica 63 (3): 375-391 - get paper here
  • Myers, C. W. 1971. Central American lizards related to Anolis pentaprion: two new species from the Cordillera de Talamanca. American Museum Novitates (2471): 1-40 [13] - get paper here
  • Nicholson, K.E. 2002. Phylogenetic analysis and a test of the current infrageneric classification of Norops (beta Anolis). Herpetological Monographs 16: 93-120 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Savage, J.M. 2002. The Amphibians and Reptiles of Costa Rica: A Herpetofauna Between Two Continents, Between Two Seas. University of Chicago Press, 934 pp. [review in Copeia 2003 (1): 205]
  • Stuart,L.C. 1955. A brief review of the Guatemalan lizards of the genus Anolis. Misc. Publ. Mus. Zool., Univ. Michigan No. 91: 1-31 - get paper here
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