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Asaccus elisae (WERNER, 1896)

IUCN Red List - Asaccus elisae - Least Concern, LC
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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Werner's Leaf-Toed Gecko 
SynonymPhyllodactylus elisae WERNER 1896: 14
Phyllodactylus eugeniae NIKOLSKY 1907
Phyllodactylus elisae — SCHMIDT 1939
Phyllodactylus eliasae — HAAS [sic] 1952
Phyllodactylus ingae EISELT 1973
Phyllodactylus elisae — ARNOLD 1977: 100
Asaccus elisae — DIXON & ANDERSON 1973
Asacus [sic] elisae — BARAN & GRUBER 1982
Asaccus elisae — KLUGE 1993
Asaccus elisae — RÖSLER 1995: 98
Asaccus elisae — PAPENFUSS et al. 2010
Asaccus elisae — NASRABADI et al. 2017 
DistributionS Turkey (Birecik, SE Anatolia, Sanliurfa: HR 33: 223)
E Iraq, SW Iran, NE Syria (Niniveh)

Type locality: ruins of Niniveh (near Mosul)

eugeniae: Iran; Type locality: Dizful and Abu-Garia, affluent to Karun River, Iran [= Dezful, Khuzestan Province, Iran, 32.38 N 48.41 E]  
Reproductionoviparous 
TypesSyntypes: NMW (4 spec.); BMNH 95.3.2.3 (1946.8.24.39)
Syntypes ZISP (also as ZIL) 10261 [6+], 10262 [6+], 10263 [4], 10270[3], 10349) [Phyllodactylus eugeniae] 
DiagnosisDiagnosis: Easily distinguishable from other Asaccus species by the strongly keeled dorsal tubercles, the length of the tubercles being equal to the length of the oval ear opening, enlarged tubercles on upper arm, and whorls of keeled and pointed tubercles and scales on the tail. Tail length 55-70 mm, thinner and much longer than snout-vent length. Body slender, about 45-50 mm; dorsum with two series of dark spots on each side of vertebral line. Male with 4 wide black bands on the tail. Lamellae adhesive pads smaller than ear opening, claws extending somewhat the digit scansors. 
CommentPhylogenetics (genus): Tamar et al. 2018, Fattahi et al. 2020 found many unnamed clades within (Iranian) Asaccus, although they did not name any of them.

Synonymy: mostly after ARNOLD & GARDNER 1994. Not listed by Engelmann et al. (1993).

Type species: Phyllodactylus elisae WERNER 1895 is the type species of the genus Asaccus DIXON & ANDERSON 1973.

Distribution: See map in SMID et al. 2014 for distribution in Iran. 
EtymologyNot given by Werner but probably his wife or daughter. 
References
  • Afrasiab, Saman R. and Sarbaz I. Mohamad 2009. A study on cave-dwelling geckos in Iraq, with the description of a new species from Saffine mountain (Reptilia: Gekkonidae). Zoology in the Middle East 47:49-56 - get paper here
  • ANANJEVA, NATALIA B.; KONSTANTIN D. MILTO, ANDREI V. BARABANOV, EVGENY A. GOLYNSKY 2020. An annotated type catalogue of amphibians and reptiles collected by Nikolay A. Zarudny in Iran and Middle Asia. Zootaxa 4722 (2): 101–128 - get paper here
  • Anderson, S.C. 1963. Amphibians and Reptiles from Iran. Proc. Cal. Acad. Sci. Ser. 4, 31(16): 417-498 - get paper here
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • Arnold E N; Gardner A S 1994. A review of the middle eastern leaf-toed geckoes (Gekkonidae: Asaccus) with descriptions of two new species from Oman. Fauna of Saudi Arabia 14: 424-441 - get paper here
  • Arnold, E. N. 1977. Little-known geckoes (Reptilia: Gekkonidae) from Arabia with descriptions of two new species from the Sultanate of Oman. The Scientific Results of the Oman Flora and Fauna Survey, 1975. Journal of Oman Studies, Special Report, (1):81—110 [1975]
  • Baran, Ibrahim; Yusuf Kumlutas; Cetin Ilgaz; Oguz Turkozan, and Aziz Avci 2003. ON Asaccus elisae (F. WERNER, 1895) (SAURIA: GEKKONIDAE) SPECIMENS COLLECTED FROM THE VICINITY OF HARRAN, SANLIURFA IN SOUTHEAST TURKEY. Russ. J. Herpetol. 10 (1): - get paper here
  • Dixon, J.R. & Anderson,S.C. 1973. A new genus and species of gecko (Sauria: Gekkonidae) from Iran and Iraq. Bull. South. Calif. Acad. Sci. 72: 155-160 [1975] - get paper here
  • Eiselt, J. 1973. Ein neuer Blattfinger-Gecko (Phyllodactylus, Sauria, Rept.) aus dem Iran und Bemerkungen zu Phyllodactylus elisae. Annalen des Naturhistorischen Museums in Wien, 77:173—179 - get paper here
  • Esterbauer, H. 1992. Asaccus elisae (Werner 1895) in Syrien. Sauria 14 (3): 31-34 - get paper here
  • Fathnia, B.; N. Rastegar Pouyani; M. Sampour; A.M. Bahrami; G. Jaafari 2009. The lizard fauna of Ilam province, Southwestern Iran. Iranian Journal of Animal Biosystematics 5 (2) - get paper here
  • Fattahi, Akbar; Nasrullah Rastegar-Pouyani, Eskandar Rastegar-Pouyani, Rasoul Karamiani, Seyyed Saeed Hosseinian Yousefkhani & Behzad Fathinia 2020. Molecular phylogeny and taxonomic evaluation of the genus Asaccus Dixon and Anderson, 1973 (Reptilia: Phyllodactylidae) in Iran. The Herpetological Journal 30: 207-214 - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Haas G. and Werner, Y. L. 1969. Lizards and snakes from Southwestern Asia, collected by Henry Field. Bull. Mus. Comp. Zool. Harvard 138: 327-406. - get paper here
  • Kamali, Kamran 2020. A Field Guide to the Reptiles and Amphibians of Iran. Edition Chimaira, Frankfurt am Main, Germany (www.chimaira.de). 574 pp.
  • Karamiani, Rasoul, Nasrullah Rastegar-Pouyani, Razieh Fattahi and Behzad Fathinia. 2013. Sexual dimorphism in leaf-toed gecko Asaccus elisae (Werner, 1895) (Sauria: Gekkonidae) from western Iran. Hamadryad 36 (2): 157-161 - get paper here
  • LEPTIEN, R., T. KOWALSKI & H.J. ZILGER 1994. Der "Große Asaccus" aus den Omanbergen, Asaccus elisae (WERNER, 1895) s. lat. Sauria 16 (1): 21-25 - get paper here
  • Leviton,A.E.; Anderson,S.C.; Adler, K.; Minton,S.A. 1992. Handbook to Middle East Amphibians and Reptiles. SSAR, Oxford, Ohio (Contr. to Herpetol. No. 8), 1-252
  • NASRABADI, REZA; NASRULLAH RASTEGAR-POUYANI, ESKANDAR RASTEGAR-POUYANI, AHMAD GHARZI 2017. A revised key to the lizards of Iran (Reptilia: Squamata: Lacertilia). Zootaxa 4227 (3): 431–443 - get paper here
  • Nikolsky, A. M. 1907. Reptiles et amphibiens recueillis (part.) M. N. A. Zarudny en Perse en 1903-1904 [in Russian and Latin]. Ann. Mus. Zool. Acad. Imp. Sci., St. Petersburg 10 [1905]: 260-301 - get paper here
  • PAPENFUSS, THEODORE J.; TODD R. JACKMAN, AARON M. BAUER, BRYAN L. STUART, MICHAEL D. ROBINSON, and JAMES F. PARHAM 2010. Phylogenetic Relationships Among Species of Southwest Asian Leaf-toed Geckos (Asaccus). Proceedings of the California Academy of Sciences, 61 (13): 587-596 - get paper here
  • Parsa, H.; H. Oraie; A. Khosravani; N. Rastegar Pouyani 2009. Systematics and Distribution of the Iranian Plateau Leaf-toed Geckos of the Genus Asaccus (Sauria: Gekkonidae). Iranian Journal of Animal Biosystematics 5 (2) - get paper here
  • Rastegar-Pouyani, N., Nilson, G. & Faizi, H. 2006. A new Species of Asaccus (Sauria: Gekkonidae) from Kurdistan Province, western Iran. Hamadryad 30: 141-150. - get paper here
  • Rastegar-Pouyani, Nasrullah 2006. Systematics of the genus Asaccus (Sauria: Gekkonidae) on the Zagros Mountains, Iran. Herpetologica Bonnensis II: 117-119 - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Rösler, Herbert 1995. Geckos der Welt - Alle Gattungen. Urania, Leipzig, 256 pp.
  • Schmidt,K.P. 1939. Reptiles and amphibians from Southwestern Asia. Publ. Field Mus. nat. Hist., zool. Ser., 24: 49-92 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, Roberto; Alberto Venchi, Giuseppe M. Carpaneto, Marco A. Bologna 2000. The Reptiles of Anatolia: a Checklist and Zoogeographical analysis. Biogeographia, 21-2000: 441-554. - get paper here
  • ŠMÍD, JIŘÍ; JIŘÍ MORAVEC, PETR KODYM, LUKÁŠ KRATOCHVÍL, SEYYED SAEED HOSSEINIAN YOUSEFKHANI, ESKANDAR RASTEGAR-POUYANI & DANIEL FRYNTA 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa 3855 (1): 001–097 - get paper here
  • Tamar, Karin; Pelagia Mitsi; Salvador Carranza 2018. Cryptic diversity revealed in the leaf‐toed gecko Asaccus montanus (Squamata, Phyllodactylidae) from the Hajar Mountains of Arabia. J Zool Syst Evol Res. 2018 ;00:1–14 - get paper here
  • Tok, C.V.; Atatür, M.K.; Taskavak, E. 1997. Taxonomic status of Asaccus elisae (F. WERNER) from Birecik, Anatolia. Israel Journal of Zoology 43: 71-77
  • Tok, Cemal Varol; Mehmet Kutsay ATATÜR & Durmus CIHAN 2002. On an Asaccus elisae(F. Werner, 1895) (Sauria: Gekkonidae) Specimen Collected from the Vicinity of Nusaybin, Mardin. Turkish Journal of Zoology 26: 315-316 - get paper here
  • Torki F., Gharzi A., Nazari-Serenje F., Javanmardi S., Heidari N., Azizpourian A., Mahdavi S., and Mahdavi-Zarkhoni M. A. 2008. Geckos of the Genera Tropiocolotes and Asaccus in the Zagros Mountains, Iran. Gekko 5 (2): 31-43
  • TORKI, F. 2010. Beschreibung eines neuen Asaccus (Reptilia: Phyllodactylidae) aus der Provinz Lorestan, Iran. Sauria 32 (4): 3-16 - get paper here
  • Torki, F. 2011. Beschreibung eines neuen Asaccus (Sauria: Phyllodactylidae) aus dem Iran. Sauria 33 (1): 51–62 - get paper here
  • Torki, F.; Ahmadzadeh, F.; Ilgaz, C.; Avci, A. & Kumlutas, Y. 2011. Description of four new Asaccus Dixon and Anderson, 1973 (Reptilia: Phyllodactylidae) from Iran and Turkey. Amphibia-Reptilia 32 (2): 185-202 - get paper here
  • UGURTAS¸ Ismail HAKKI; HIKMET SAMI YILDIRIMHAN & Murat Sevinc 2007. Distribution of the Gekkonidae Species in Southeast Anatolia, Turkey, and New Localities. Turkish Journal of Zoology 31: 137-141 - get paper here
  • Vogel, G. 1989. Das Portrait: Asaccus elisae (WERNER). Sauria 11 (3): 1-2 - get paper here
  • Werner,F. 1896. Über eine Sammlung von Reptilien aus Persien, Mesopotamien und Arabien. Verh. Zool.-Bot. Ges. Wien 45: 13-22 [1895] - get paper here
 
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