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Bronchocela hayeki (MÜLLER, 1928)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Sumatra Bloodsucker 
SynonymCalotes hayeki MÜLLER 1928
Calotes hayeki — BRONGERSMA 1931
Calotes hayeki — WERMUTH 1967: 38
Bronchocela hayeki — MOODY 1980
Bronchocela hayeki — MANTHEY & GROSSMANN 1997: 161
Bronchocela hayeki — TEYNIÉ et al. 2010
Bronchocella hayeki — NUGRAHA et al. 2020 (in error) 
DistributionIndonesia (N Sumatra)

Type locality: Hochfläche von Brastag, am Sibayak, Nordost-Sumatra (elevation 1400 m). Neotype locality: near Berastagi 3°11’ N, 98° 31’ E, “Fuss des Sibayak” about 1450 m a.s.l., Sumatera Utara.  
TypesNeotype. ZMB 55931 near Berastagi 3°11’ N, 98° 31’ E, “Fuss des Sibayak” about 1450 m a.s.l., Sumatera Utara, leg. Ulrich Manthey, April 1996, adult male (designated by Hallermann 2005). Holotype formerly ZSM 3/1928, destroyed fide Glaw, cited in Hallermann 2005.
DiagnosisDiagnosis. Bronchocela hayeki can be distinguished from its sympatric congener B. cristatella by the presence of larger gular scales and larger ventral scales in 8 rows (vs. 10 – 12 in cristatella) and crescent-shaped erect nuchals as long as or longer than diameter of orbit in males and a dark brown tympanum and dark colored orbit; from jubata and orlovi with similar nuchal scales and shape of gular sac by higher number of scales around midbody (58 – 75 vs. 43 – 55), from celebensis by a larger diameter of tympanum (more than half diameterof orbit). It differs from B. marmorata by more dorsal scale rows directed upward (7 – 9 vs. 1 – 2 in marmorata, jubata, orlovi, and 3 – 4 in celebensis). It differs from B. danieli, smaragdina and Bronchocela sp. nov. by fourth finger not longer than fifth toe. 
EtymologyNamed after Hans von Hayek (1869-1940), an Austrian artist who moved to Munich (1891) and traveled widely within Europe and Asia. 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • GRISMER, L. LEE; P.L. WOOD, JR., CHEOL HAENG LEE, EVAN S. H. QUAH, SHAHRUL ANUAR, EHWAN NGADI & JACK W. SITES, JR. 2015. An integrative taxonomic review of the agamid genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with descriptions of new montane and insular endemics. Zootaxa 3948 (1): 001–023 - get paper here
  • Hallermann, J. 2005. A taxonomic review of the genus Bronchocela (Squamata: Agamidae) with description of a new species from Vietnam. Russ. J. Herpetol. 12 (3): 167-182 - get paper here
  • Hallermann, J. 2005. The bizarre arboreal agamids. Reptilia (GB) (42): 8-15 - get paper here
  • Hallermann, J. 2005. Mit Hörnern, Kämmen und Gleithäuten - die bizarren Baumagamen. Reptilia (Münster) 10 (51): 18-25 - get paper here
  • Malkmus,R. & Kunkel, M. 1992. Amphibien und Reptilien aus dem Leuser-Nationalpark/Nordsumatra. Herpetofauna 14 (76): 27 - get paper here
  • Manthey U 2008. Agamid lizards of Southern Asia, Draconinae 1. Terralog 7, 160 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Moody,S.M. 1980. Phylogenetic and historical biogeographical relationships of the genera in the family Agamidae (Reptilia: Lacertilia). PhD thesis, Univ. Michigan, Ann Arbor, 373 pp.
  • Müller, L. 1928. Herpetologische Mitteilungen II. Ein neuer Calotes von Sumatra. Zool. Anz. 77: 67-69
  • Nugraha, F. A. D., Selaras, G. H., & Satria, R. 2020. Preliminary Checklist of Herpetofauna of Mount Sago Along the Hiking Trail in the Dry Season. International Conference on Biology, Sciences and Education (ICoBioSE 2019) (pp. 51-55). Atlantis Press - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
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