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Bronchocela rubrigularis HALLERMANN, 2009

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Red-throated bloodsucker 
SynonymBronchocela rubrigularis HALLERMANN 2009
Bronchocela rubrigularis — AMARASINGHE et al. 2022 
DistributionIndia (Nicobar Islands)

Type locality: Trinkat Island, central Nicobar Islands  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZMH R09271 (Fieldnumber Tri 25) (see Fig. 2 in Hallermann 2009). An adult male collected by S.P. Vijayakumar on Nicobar in 2004. Other specimens: ZSI 
DiagnosisDiagnosis: A relative robust species with a SVL of 81–106 mm, and a long tail (280–377 mm (336–378 % of SVL). Tympanum large, more than half diameter of orbit, often dusky coloured, Ratio tympanum /orbit 0.74–0.89. Head covered above with small keeled scales, up to two slightly enlarged scales between orbit and tympanum. 7–10 supralabials and infralabials on each side. Canthus rostralis sharp, two small erect, compressed scales behind supraciliary edge. Nuchal crest formed by 7–10 lanceolate erect scales, bordered laterally by smaller erect scales, longest little longer than diameter of orbit. Dorsal crest smaller than nuchal crest, continues nuchal crest by a small gap. Mental wider than high, three postmentals, gular sac small in males, gular region covered with little enlarged keeled scales, smaller than ventrals.
Body scales mucronate, keeled, homogenous in 52–58 midbody scales. 1–3 uppermost scale rows next to dorsal crest pointing upward, 4–5 rows directed parallel, others scale rows directed downwards. Ventrals about two times larger than dorsals, strongly keeled. It can be distinguished from other congeners of the Nicobar Islands: from B. cristatella by a larger nuchal crest, only 1–4 upper scale rows pointing upwards (versus 5–10), and a red gular patch (white in alcohol) in males; from B. danieli by a larger tympanum (versus only half diameter of orbit) , longer fifth toe than fourth finger (versus fifth toe smaller than fourth finger) and only two times larger ventrals (versus five times larger) (Hallermann 2009) 
Comment 
EtymologyThe species is named after its red gular (latin rubber = red, gula (feminin) = gular) 
References
  • AMARASINGHE, ; A.A. THASUN; IVAN INEICH, AWAL RIYANTO, JAKOB HALLERMANN, NOVIAR ANDAYANI, A. ABINAWANTO, JATNA SUPRIATNA 2022. Taxonomy and distribution of a common arboreal lizard, Bronchocela jubata Duméril & Bibron, 1837 (Reptilia: Agamidae), with designation of its lectotype from Java, Indonesia. Zootaxa 5150 (1): 065–082 - get paper here
  • CHANDRAMOULI, S., ADHIKARI, O. D., AMARASINGHE, A. T., & ABINAWANTO, A. 2023. A review of the genus Bronchocela Kaup, 1827 (Reptilia: Agamidae) in the Nicobar Archipelago with the description of two new species. Zootaxa 5254 (4): 493-516 - get paper here
  • GRISMER, L. LEE; P.L. WOOD, JR., CHEOL HAENG LEE, EVAN S. H. QUAH, SHAHRUL ANUAR, EHWAN NGADI & JACK W. SITES, JR. 2015. An integrative taxonomic review of the agamid genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with descriptions of new montane and insular endemics. Zootaxa 3948 (1): 001–023 - get paper here
  • Hallermann, J. 2009. A new species of Bronchocela (Squamata: Agamidae) from Nicobar Island. Bonner zoologische Beiträge 56 (4): 279–284 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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