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Crossobamon eversmanni (WIEGMANN, 1834)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
SubspeciesCrossobamon eversmanni eversmanni (WIEGMANN 1834)
Crossobamon eversmanni lumsdenii (BOULENGER 1887) 
Common NamesE: Comb-toed Gecko
G: Kammzehengecko 
SynonymGymnodactylus eversmanni WIEGMANN 1834: 19
Crossobamon atropunctatus LICHTENSTEIN & VON MARTENS 1856
Gymnodactylus atropunctatus — BOULENGER 1885: 22
Crossobamon eversmanni — WERMUTH 1965: 22
Gymnodactylus eversmanni — BAUER & GÜNTHER 1991
Crossobamon eversmanni — DAS 1996: 40
Crossobamon eversmanni — SZCZERBAK 2003
Grossobamon eversmanni — ZAVYALOV et al. 2006: 38 (in error)
Crossobamon eversmanni — ŠMÍD et al. 2014
Crossobamon eversmanni — RÖSLER 2018: 43

Crossobamon eversmanni lumsdenii (BOULENGER 1887)
Stenodactylus lumsdenii BOULENGER 1887: 479
Stenodactylus maynardi SMITH 1933: 18
Crossobamon maynardi — KLUGE 1967: 23
Crossobamon eversmanni lumsdenii — SZCZERBAK & GOLUBEV 1986
Crossobamon eversmanni maynardi — KLUGE 1991: 6
Crossobamon eversmanni lumsdenii — RÖSLER 2000: 64
Crossobamon eversmanni — KHAN (pers. comm.)
Crossobamon lumsdenii — KHAN 2003
Crossobamon maynardi — KHAN 2003
Crossobamon lumsdeni — KHAN 2004
Crossobamon maynardi — KHAN 2004
Crossobamon maynardi — KRYSKO et al. 2007
Crossobamon eversmanni lumsdenii — KHANI et al. 2013 
DistributionNW Afghanistan, N Iran, Pakistan, Turkmenistan, Uzbekistan, S Kazakhstan, S Tajikistan, Kyrgyzstan

lumsdenii: Pakistan

maynardi: Pakistan; Type locality: “Baluchistan near the Afghan border”

Type locality: Kyzyl Kum Desert
Type locality: Agetma, Zentral-Asien (fide WERMUTH 1965)  
Reproductionoviparous 
TypesHolotype: ZMB 435
Holotype: ZMB 435 [atropunctatus] (see BAUER & GÜNTHER 1991)
Syntypes: ZSI 139945, BMNH 1946.8.23.51 [lumsdenii]
Syntypes: ZSI 13944, BMNH originally 1931.6-14-1 [maynardi] 
DiagnosisDiagnosis: Back with small dark dots, irregularly arranged or fonning 10-12 more or less continuous, vermiculate rows; dark stripes that extend from nostrils over eye and above ear usually end on flanks in first third of body; tail with up to 30 transverse bars, which may coalesce to form longitudinal stripes on sides; 5-11 gular scales in contact with mental (Szczerbak and Golubev 1986:49; 1996:48, Anderson 1999: 147). 
CommentSynonymy: Given that lumsdenii and maynardi are considered as full species by KHAN (2003), evermanni s. str. does not occur in Pakistan according to KHAN (pers. comm.). However, KRYSKO et al. (2007) state the the status of C. lumsdenii is questionable.

Type species: Gymnodactylus eversmanni WIEGMANN 1834: 19 is the type species of the genus Crossobamon BOETTGER 1888 (nomen novum pro Ptenodactylus STRAUCH 1887 non Ptenodactylus GRAY 1845).

Distribution: See map in SMID et al. 2014 for distribution in Iran. 
EtymologyNamed after Eduard Eversmann (1794-1860), German physician and naturalist. 
References
  • Ananjeva, NB, Orlov NL; Khalikov RG; Darevsky IS; Ryabov IS & Barabanov AV 2006. The Reptiles of North Eurasia. Taxonomic Diversity, Distribution, Conservation Status [this comprises the territory of the former Soviet Union and Mongolia]. Pensoft Series Faunistica 47, 250 pp.
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • Bauer, A.M. & Günther,R. 1991. An annotated type catalogue of the geckos (Reptilia: Gekkonidae) in the Zoological Museum , Berlin. Mitt. Zool. Mus. Berlin 67: 279-310 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bondarenko D. A., Ergashev U. H. 2022. Reptiles of the Southwestern Tajikistan desert plains: Spatial distribution, population density and communities structure. [in Russian] Current Studies in Herpetology 22 (1-2): 17 - get paper here
  • Bondarenko, D.A. & Evgeny A. Peregontsev 2009. Peculiarities of landscape distribution of the Central Asian Tortoise, Agrionemys horsfieldii, in Iran. Zoology in the Middle East 48: 49–62 - get paper here
  • Boulenger, G. A. 1887. Catalogue of the lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575 pp. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Brushko, Z.K. 1995. Lizards of desert regions of Kazakhstan. [in Russian] Kazakhstan National Academy of Sciences, Institute of Zoology, Almaty, 232 pp.
  • Davletbakov, A. T.; L. A. Kustareva, D. A. Milko, A. N. Ostastshenko, S. S. Sagymbaev, S. A. Toropov, N. V. Trotchenko 2015. Genetical Fund Cadastre of Kyrgyz Republic. Vol. IV. Phylum Chordata (chordates). Bishkek, 128 p. + XXXVI color tables - get paper here
  • Kamali, Kamran 2020. A Field Guide to the Reptiles and Amphibians of Iran. Edition Chimaira, Frankfurt am Main, Germany (www.chimaira.de). 574 pp.
  • Khan, M.S. 2003. Anmerkungen zur Morphologie, Verbreitung und den Habitatpräferenzen einiger pakistanischer Geckos. Sauria 25 (3): 35-47 [erratum in 25 (4): 27] - get paper here
  • Khani, Ali; Masoud Yousefi, Sayyad Sheykhi, Ali Sheykhi, Seyyed Saeed Hosseinian Yousefkhani and Eskandar Rastegar-Pouyani 2013. New record of the gecko Crossobamon eversmanni lumsdeni (Boulenger, 1887) from Parvand Protected Area, Khorasan Razavi province, in eastern Iran. Herpetology Notes 6: 101-102
  • Kluge, A. G. 1967. Higher taxonomic categories of gekkonid lizards and their evolution. Bull. Amer. Mus. Nat. Hist., 135:1-60 - get paper here
  • Krysko, K. L.; Rehman, H. & Auffenberg, K. 2007. A new species of Cyrtopodion (Gekkonidae: Gekkoninae) from Pakistan. Herpetologica 63 (1): 100-113 - get paper here
  • Leviton, Alan E. 1959. Report on a collection of reptiles from Afghanistan. Proc. Cal. Acad. Sci. 29 (12): 445-463 - get paper here
  • Lichtenstein, H., and E. von Martens. 1856. Nomenclator reptilium et amphibiorum Musei Zoologici Berolinensis. Namenverzeichniss der in der zoologischen Sammlung der Königlichen Universität zu Berlin aufgestellten Arten von Reptilien und Amphibien nach ihren Ordnungen, Familien und Gattungen. Königliche Akademie der Wissenschaften, Berlin / Buchdruckerei der Königl. Akad. der Wiss., iv + 48 pp. - get paper here
  • Nikolsky, A. M. 1897. Les reptiles, amphibiens et poissons recueillis (part.) Mr. N. Zaroudny dans la Perse orientale [in Russian]. Annuaire Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg 2: 306-348 - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Rösler, H. 1979. Eine siamesische Zwillingsbildung bei der Gecko-Species Crossobamon eversmanni (Reptilia: Sauria: Gekkonidae). Salamandra 15 (2): 108-110 - get paper here
  • Rösler, H. 1990. Crossobamon eversmanni (WIEGMANN, 1834). Sauria 12 Suppl.: 179–182 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, H. 2018. Haftbar -- Die einzigartige Welt der Geckos. Begleitheft zur Ausstellung im Naturkundemuseum Erfurt Mai-Aug 2018, 96 pp.
  • Rösler, Herbert 1997. Paläarktische Geckos (Reptilia: Gekkota) Teil I: Historischer Überblick, Geographie und Klima, Verbreitung. Sauria 19 (1): 17-28 - get paper here
  • Rösler, Herbert 2017. Gecko-Chorologie (Squamata: Gekkota). Gekkota (4): 1-160
  • SAFAEI-MAHROO, BARBOD; HANYEH GHAFFARI, & STEVEN C. ANDERSON 2016. A new genus and species of gekkonid lizard (Squamata: Gekkota: Gekkonidae) from Hormozgan Province with a revised key to gekkonid genera of Iran. Zootaxa 4109 (4): 428–444 - get paper here
  • Shcherbak, N. N. [= Szczerbak], and M. L. Golubev 1996. Gecko fauna of the USSR and contiguous regions. SSAR [Russian original published in 1986]. [book review in Salamandra 36 (2): 138]
  • Shestopal A., Amanov A., Ovezov T. 2021. An overview of reptiles from the Kaplankyr Nature Reserve and adjacent territories. [in Russian] Proc. Inst. Zool., Kazakhstan, pp. 105-126 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • ŠMÍD, JIŘÍ; JIŘÍ MORAVEC, PETR KODYM, LUKÁŠ KRATOCHVÍL, SEYYED SAEED HOSSEINIAN YOUSEFKHANI, ESKANDAR RASTEGAR-POUYANI & DANIEL FRYNTA 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa 3855 (1): 001–097 - get paper here
  • Smith, M.A. 1933. Remarks on some old world geckoes. Records of the Indian Museum 35: 9-19.
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp. - get paper here
  • Wiegmann,A.F.A. 1834. Herpetologia Mexicana, seu descriptio amphibiorum novae hispaniae , quae itineribus comitis de Sack, Ferdinandi Deppe et Chr. Guil. Schiede im Museum Zoologicum Berolinense Pervenerunt. Pars prima, saurorum species. Berlin, Lüderitz, iv + 54 pp. - get paper here
 
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