Ctenophorus clayi (STORR, 1967)

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Higher Taxa	Agamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards)
Subspecies
Common Names	E: Black-shouldered Ground-dragon
Synonym	Amphibolurus clayi STORR 1967: 24 Amphibolurus clayi — COGGER 1983 Ctenophorus raffertyi WELLS & WELLINGTON 1985: 17 (fide SHEA 1999) Ctenophorus clayi — GREER 1991 Ctenophorus clayi — COGGER 2000: 311 Ctenophorus clayi — WILSON & SWAN 2010 Ctenophorus clayi — CHAPPLE et al. 2019: 58
Distribution	Australia (Northern Territory, South Australia, Western Australia) Type locality: 3 mi S of Learmonth, in 22° 16’ S, 114'06’ E, Western Australia
Reproduction	oviparous
Types	Holotype: WAM R14462
Diagnosis
Comment	Lifestyle: burrowing. Abundance: common (Pianka 2011) Diet: a large percentage of the diet of this species (10-30%) consists of ants (Lucas et al. 2023).
Etymology	Named after Brian T. Clay (1950-2004), a forestry official in East Africa who moved to Australia (1960), where he worked as a field assistant for the Zoology Department, University of Western Australia. Clay helped to collect the holotype.
References	Anonymous 1967. Type Specimens in the Western Australian Museum (Part 8). Annual Report, Western Australian Museum, 1966-67, Perth, pp. 59-60 Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here Chapple, David G.; Reid Tingley, Nicola J. Mitchell, Stewart L. Macdonald, J. Scott Keogh, Glenn M. Shea, Philip Bowles, Neil A. Cox, John C. Z. Woinarski 2019. The Action Plan for Australian Lizards and Snakes 2017. CSIRO, 663 pp. DOI: 10.1071/9781486309474 - get paper here Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp. Doughty, P.; Maryan, B.; Melville, J. & Austin, J. 2007. A new species of Ctenophorus (Lacertlia: Agamidae) from Lake Disappointment, Western Australia. Herpetologica 63 (1): 72-86 - get paper here Ellis, Ryan J. 2019. An annotated type catalogue of the dragon lizards (Reptilia: Squamata: Agamidae) in the collection of the Western Australian Museum. Records of the Western Australian Museum 34: 115–132 - get paper here Greer A E 1991. Limb reduction in Squamates: identification of the lineages and discussion of the trends. Journal of Herpetology 25 (2): 166-173 - get paper here Greer, A.E. 1987. Observations on the osteology and natural history of the agamid lizard Ctenophorus clayi. Western Australian Naturalist 17 (1): 5-7 - get paper here Lucas B Q Cavalcanti, Gabriel C Costa, Guarino R Colli, Eric R Pianka, Laurie J Vitt, Daniel O Mesquita, 2023. Myrmecophagy in lizards: evolutionary and ecological implications. Zoological Journal of the Linnean Society, 2023;, zlad175, - get paper here Melville, Jane; Schulte, James A.; Larson, Allan 2001. A molecular phylogenetic study of ecological diversification in the Australian lizard genus Ctenophorus. Journal of Experimental Zoology: Molecular and Developmental Evolution 291(4): 339-353 - get paper here Pianka, E. R. 1969. Habitat specificity, speciation, and species density in Australian desert lizards. Ecology 50 (3): 498-502 - get paper here Storr, G. M. 1966. The Amphibolurus reticulatus species-group (Lacertilia: Agamidae) in Western Australia. J R. Soc. West. Aust. 49: 17-25 Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
External links	IUCN Red List National Center for Biotechnology Information http://www.amonline.net.au/herpetology/research/pdf/agamidae.pdf http://www.tropicalherping.com/publications/books/mindo/riama-oculata.html http://www.arod.com.au/arod/reptilia/Squamata/Agamidae/Ctenophorus/clayi http://www.arod.com.au/arod/reptilia/Squamata/Elapidae/Demansia/shinei Google images

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