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Ctenophorus scutulatus (STIRLING & ZIETZ, 1893)

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Lozenge-marked Bicycle-dragon, Lozenge-Marked Dragon 
SynonymAmphibolurus scutulatus STIRLING & ZIETZ 1893: 165
Amphibolurus websteri BOULENGER 1904: 414
Amphibolurus holsti ROSÉN 1905
Amphibolurus scutulatus — FRY 1914: 182
Amphibolurus scutulatus — LICHT et al. 1966
Amphibolurus scutulatus — COGGER 1983
Licentia scutulata — WELLS & WELLINGTON 1985: 18
Licentia websteri — WELLS & WELLINGTON 1985: 18
Ctenophorus scutulatus — COGGER 2000: 326
Ctenophorus scutulatus — WILSON & SWAN 2010
Ctenophorus scutulatus — CHAPPLE et al. 2019: 70 
DistributionAustralia (South Australia, Western Australia)

Type locality: between Fraser Range and Queen Victoria Springs, W. A.  
Reproductionoviparous 
TypesLectotype: SAMA R3024 (the larger of the 2 specimens under this number), designation by Wells & Wellington (1984
Syntypes: BMNH 1946. 9. 4. 31-32, from Coolgardie District, 400 mi inland of Perth, W. A. [Amphibolurus websteri]
Holotype: MZLU (was ZMLU) Cat. Typ. Nr. 284, from W. A. [Amphibolurus holsti] 
Diagnosis 
CommentSynonymy after COGGER 1983.

Habitat: shrubs, hummock grass.

Diet: a large percentage of the diet of this species (10-30%) consists of ants (Lucas et al. 2023). 
EtymologyNamed after the Latin scutula, "a small plate," and -atus, "provided with," referring to the perceived smaller scales than in other populations. 
References
  • Boulenger, G. A. 1904. Description of a new lizard from Western Australia. Ann. Mag. Nat. Hist. (7) 14: 414-415 - get paper here
  • Chapple, David G.; Reid Tingley, Nicola J. Mitchell, Stewart L. Macdonald, J. Scott Keogh, Glenn M. Shea, Philip Bowles, Neil A. Cox, John C. Z. Woinarski 2019. The Action Plan for Australian Lizards and Snakes 2017. CSIRO, 663 pp. DOI: 10.1071/9781486309474 - get paper here
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Doughty, P.; Maryan, B.; Melville, J. & Austin, J. 2007. A new species of Ctenophorus (Lacertlia: Agamidae) from Lake Disappointment, Western Australia. Herpetologica 63 (1): 72-86 - get paper here
  • Fry, D. B. 1914. On a collection of reptiles and batrachians from Western Australia. Rec. West. Austral. Mus. 1:174-210 - get paper here
  • Glauert, L. 1959. Herpetological Miscellanea. XI. Dragon Lizards of the Genus Amphibolurus. Western Australian Naturalist 7 (2): 42-51 - get paper here
  • Houston, T. F. 1976. Vertebrate type-specimens in the South Australian Museum. III. Reptiles. [type catalogue] Rec. South Austral. Mus. 17: 181-187 - get paper here
  • Licht, P., W.R. Dawson, V.H. Shoemaker & A.R. Main 1966. Observations on the thermal relations of Western Australian lizards Copeia 1966 (1): 97-110. - get paper here
  • Lucas B Q Cavalcanti, Gabriel C Costa, Guarino R Colli, Eric R Pianka, Laurie J Vitt, Daniel O Mesquita, 2023. Myrmecophagy in lizards: evolutionary and ecological implications. Zoological Journal of the Linnean Society, 2023;, zlad175, - get paper here
  • Melville, Jane; Schulte, James A.; Larson, Allan 2001. A molecular phylogenetic study of ecological diversification in the Australian lizard genus Ctenophorus. Journal of Experimental Zoology: Molecular and Developmental Evolution 291(4): 339-353 - get paper here
  • Pianka, E. R. 1969. Habitat specificity, speciation, and species density in Australian desert lizards. Ecology 50 (3): 498-502 - get paper here
  • Pianka, E. R. 1971. Notes on the biology of Amphibolurus cristatus and Amphibolurus scutulatus. Western Australian Naturalist 12: 36-41 - get paper here
  • Rosen, N. 1905. List of the lizards in the Zoological Museum of Lund, with descriptions of new species. Ann. Mag. nat. Hist. (7) 16:129-142 - get paper here
  • Stirling, E. C., and A. Zietz. 1893. Scientific results of the Elder Exploring Expedition. Vertebrata. Mammalia, Reptilia. Transactions of the Royal Society of South Australia, 16:154-176. - get paper here
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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