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Cubatyphlops anousius (THOMAS & HEDGES, 2007)

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Higher TaxaTyphlopidae (Typhlopinae), Typhlopoidea, Serpentes, Squamata (snakes)
Common NamesE: Cuban Pallid Blindsnake 
SynonymTyphlops anousius THOMAS & HEDGES 2007: 20
Cubatyphlops anousius — HEDGES et al. 2014
Typhlops anousius — PYRON & WALLACH 2014
Cubatyphlops anousius — NAGY et al. 2015
Typhlops anousius — WALLACH et al. 2014: 758 
DistributionCuba (Guantánamo Province)

Type locality: 3.5 km E Tortuguilla, Guantánamo Province, Cuba, 10 meters elevation.  
TypesHolotype: MNHNCu 4550 (field tag number 191295), on 28 June 1990 by S. Blair Hedges, Richard Thomas, and Daniel McCallister. 
DiagnosisDiagnosis: An apparently small species of Typhlops of the T. biminiensis species group distinguished from other members of the group by a lack of pigment (except for the eyes). From T. arator it can be distinguished by a relatively smaller and differently shaped rostral (Figs. 8, 10A in THOMAS & HEDGES 2007) and fewer dorsal scales (465–513 versus 578–579). The rostral of T. anousius is narrower dorsally (RW1) and at the labial border (RW3) than that of its closest relative, T. perimychus (Fig. 10E); RW1/RL1 = 0.74–0.77 versus 0.79–1.05 in T. perimychus. Typhlops anousius is the most extreme all the species in the width of the upper arm of the anterior nasal: ANTNAS/RW1 = 0.40–0.64 versus 0.34–0.39 in T. perimychus. In its narrow rostral T. anousius is somewhat similar to T. epactius, although its rostral is narrower (RW1/RL1, 0.74–0.77 versus 0.78–0.85) and straighter sided. Additionally, T. anousius differs from T. epactius in its lack of pigment with the latter having normally pigmented scale rows 9–11 (neck-midbody), its shorter tail (TL/TA 55–56 versus 40–48) and in being somewhat heavier bodied (TL/MBD 45–55 versus 55–58), and in having 24 scale rows (reducing at 2% TL) rather than 22 (no reduction). 
EtymologyFrom the Greek, anousios, meaning without substance, ghostly, in reference to its lack of pigment. 
  • DOMÍNGUEZ, MICHEL & LUIS V. MORENO 2009. Taxonomy of the Cuban blind snakes (Scolecophidia, Typhlopidae), with the description of a new large species. Zootaxa 2028: 59-66 - get paper here
  • Hedges, S.B., Marion, A.B., Lipp, K.M., Marin, J. & Vidal, N. 2014. A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology 49: 1–61 - get paper here
  • Kornilios, P.; S. Giokas, P. Lymberakis, R. Sindaco 2013. Phylogenetic position, origin and biogeography of Palearctic and Socotran blind-snakes (Serpentes: Typhlopidae). Molecular Phylogenetics and Evolution 68 (1): 35–41 - get paper here
  • NAGY, ZOLTÁN T.; ANGELA B. MARION, FRANK GLAW, AURÉLIEN MIRALLES,<br />JOACHIM NOPPER, MIGUEL VENCES & S. BLAIR HEDGES 2015. Molecular systematics and undescribed diversity of Madagascan scolecophidian snakes (Squamata: Serpentes). Zootaxa 4040 (1): 031–047 - get paper here
  • Pyron, R.A. & Wallach, V. 2014. Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence. Zootaxa 3829 (1): 001–081 - get paper here
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • THOMAS R. & S.B. HEDGES 2007. Eleven new species of snakes of the genus Typhlops (Serpentes: Typhlopidae) from Hispaniola and Cuba. Zootaxa 1400: 1-26 - get paper here
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