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Cyrtodactylus astrum GRISMER, WOOD, QUAH, ANUAR, MUIN, SUMONTHA, AHMAD, BAUER, WANGKULANGKUL, GRISMER & PAUWELS, 2012

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Starry Bent-toed Gecko 
SynonymCyrtodactylus astrum GRISMER, WOOD, QUAH, ANUAR, MUIN, SUMONTHA, AHMAD, BAUER, WANGKULANGKUL, GRISMER & PAUWELS 2012
Cyrtodactylus astrum — GRISMER & QUAH 2019 
DistributionPeninsular Malaysia (from at least La-ngu District, Satun Province, Thailand south to Kuala Perlis, Perlis)

Type locality: Wang Kelian, Perlis, Peninsular Malaysia (06°41.805 N, 100°10.751 E) at 150 meters elevation.  
Reproductionoviparous (manual and phylogenetic imputation, fide Zimin et al. 2022) 
TypesHolotype: ZRC 2.6962, adult male, collected on 24 March 2011 by L. L. Grismer, S. Anuar, E. S. H. Quah, Chan, K. O., and M. A. Muin. Paratypes. ZRC 2.6963, LSUHC 8810 and 8816 (adult male, female, and male, respectively) collected on 3 March 2008 by L. L. Grismer and Chan K. O. from Gua Kelam, Perlis, Peninsular Malaysia and an adult female (ZRC 2.6964) and adult male (ZRC 2.6965) collected on 11 October 2009 and 21 May 2010, respectively, by E. S. H. Quah, S. Anuar, and M. A. Muin from Gua Wang Burma, Perlis, Peninsular Malaysia (06°42.005 N, 100°11.878 E) at 180 meters in elevation. 
DiagnosisDiagnosis: Adult males reaching 108.3 mm SVL, adult females reaching 104.8 mm SVL; 10–12 supralabials, 9–12 infralabials; tubercles of dorsum small to moderately large with several intervening smaller tubercles; no tubercles on ventral surfaces of forelimbs, gular region, or in ventrolateral body fold; 40–57 paravertebral tubercles; 20–29 longitudinal rows of dorsal tubercles; 31–46 rows of ventral scales; 20–24 subdigital lamellae on fourth toe; 31–38 femoro-precloacal pores in males; dorsum bearing a scattered pattern of white tubercles; four dark body bands in adults with lightened centers and light colored tubercles; band to interspace ratio 1.00–2.00; 13 or 14 dark caudal bands on original tail; white caudal bands in adults heavily infused with dark pigmentation, nearly indiscernible; and posterior portion of tail in hatchlings and juveniles white. These characters are scored across all species of the Cyrtodactylus pulchellus complex in Table 6 [GRISMER et al. 2012]. 
CommentSpecimens: N=11 in original description.

Habitat: saxicolous, lowland forests associated with karst formations. 
EtymologyThe specific epithet astrum is Latin, meaning “a star” and is in reference to the starry pattern of the white dorsal tubercles. 
References
  • GOLDBERG, S., BURSEY, C.R. & GRISMER, L.L. 2020. Endoparasiten beim Stern-Bogenfingergecko Cyrtodactylus astrum von der Malaiischen Halbinsel. Sauria 42 (2): 59-60
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L.; P. L. WOOD, JR., SHAHRUL ANUAR, H. R. DAVIS, A. J. COBOS & M. L. MURDOCH 2016. A new species of karst forest Bent-toed Gecko (genus Cyrtodactylus Gray) not yet threatened by foreign cement companies and a summary of Peninsular Malaysia’s endemic karst forest herpetofauna and the need for its conservation. Zootaxa 4061 (1): 001–017 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR, SHAHRUL ANUAR, MARTA S. GRISMER, EVAN S. H. QUAH, MATTHEW L. MURDOCH, MOHD ABDUL MUIN, HAYDEN R. DAVIS, CÉSAR AGUILAR, RANDY KLABACKA, ANTHONY J. COBOS, ANCHALEE AOWPHOL, JACK W. SITES, JR 2016. Two new Bent-toed Geckos of the Cyrtodactylus pulchellus complex from Peninsular Malaysia and multiple instances of convergent adaptation to limestone forest ecosystems. Zootaxa 4105 (5): 401-429 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR., EVAN S. H. QUAH, SHAHRUL ANUAR, MOHD. ABDUL MUIN, MONTRI SUMONTHA, NORHAYATI AHMAD, AARON M. BAUER, SANSAREEYA WANGKULANGKUL, JESSE L. GRISMER & OLIVIER S. G. PAUWELS 2012. A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa 3520: 1–55
  • Grismer, L.L.; PERRY L. WOOD, SHAHRUL ANUAR, EVAN S. H. QUAH, MOHD ABDUL MUIN, MAKETAB MOHAMED, CHAN KIN ONN, ALEXANDRA X. SUMARLI, ARIEL I. LOREDO, HEATHER M. HEINZ 2014. The phylogenetic relationships of three new species of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from poorly explored regions in northeastern Peninsular Malaysia. Zootaxa 3786 (3): 359–381 - get paper here
  • LUU, VINH QUANG; THOMAS CALAME, Truong QUang Nguyen, MICHAEL BONKOWSKI & THOMAS ZIEGLER 2015. A new species of Cyrtodactylus (Squamata: Gekkonidae) from the limestone forest of Khammouane Province, central Laos. Zootaxa 4058 (3): 388–402 - get paper here
  • LUU, VINH QUANG; Truong QUang Nguyen, MINH DUC LE, MICHAEL BONKOWSKI, THOMAS ZIEGLER 2016. A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa 4079 (1): 087–102 - get paper here
  • LUU, VINH QUANG;THOMAS CALAME, MICHAEL BONKOWSKI, TRUONG QUANG NGUYEN & THOMAS ZIEGLER 2014. A new species of Cyrtodactylus (Squamata: Gekkonidae) from Khammouane Province, Laos. Zootaxa 3760 (1): 054–066 - get paper here
  • QUAH, EVAN S. H.; L. LEE GRISMER, PERRY L. WOOD, JR. & SHAHRUL ANUAR MOHD SAH 2019. The discovery and description of a new species of Bent-toed Gecko of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa 4668 (1): 051–075 - get paper here
  • Termprayoon, K., Rujirawan, A., Grismer, L. L., Wood Jr, P. L., & Aowphol, A. 2021. Taxonomic reassessment and phylogenetic placement of Cyrtodactylus phuketensis (Reptilia, Gekkonidae) based on morphological and molecular evidence. ZooKeys, 1040, 91 - get paper here
  • WOOD JR, PERRY L.; L. LEE GRISMER, MOHD ABDUL MUIN, SHAHRUL ANUAR, JAMIE R. OAKS 2020. A new potentially endangered limestone-associated Bent-toed Gecko of the Cyrtodactylus pulchellus (Squamata: Gekkonidae) complex from northern Peninsular Malaysia. Zootaxa 4751 (3): 437–460 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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