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Cyrtodactylus evanquahi WOOD, GRISMER, MUIN, ANUAR, OAKS & SITES, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Evan Quah’s banded Bent-toed Gecko
Malay: Cicak Jari-bengkok evan Quah 
SynonymCyrtodactylus evanquahi WOOD, GRISMER, MUIN, ANUAR, OAKS & SITES 2020

DistributionPeninsular Malaysia (Kedah)

Type locality: Gunung Baling, Kedah, Peninsular Malaysia (5.684989 N, 100.912590 E; 226 m elevation.  
TypesHolotype: BYU 53435, Adult male, collected on 13 August 2016 by Perry L. Wood, Jr. and Evan S. H. Quah.
Paratypes. Adult female, BYU 53436 and juvenile BYU 53437 collected on 14 August 2016 by Evan S. H. Quah and Perry L. Wood, Jr. both specimens bear the same collection data as the holotype. 
DiagnosisDiagnosis. Cyrtodactylus evanquahi sp. nov. can be differentiated from all other species of Cyrtodactylus by having a combination of the following characters: maximum SVL of approximately 96 mm; nine or 10 supralabials; nine or 10 infralabials; prominent tuberculation on body; no tubercles on ventral surface of forelimbs, gular region, in ventrolateral body folds, or anterior one-third of tail; 31–34 paravertebral tubercles; 18–23 longitudinal tubercle rows; 29–33 ventral scales; 22 or 23 subdigital lamellae on fourth toe; 32–36 femoro-precloacal pores; shallow precloacal groove in males; six or seven dark dorsal body bands; body bands much narrower than interspaces; faint rostral chevron; body bands and nuchal loop edged with a thin white, tubercle-bearing line; dorsum lacking scattered pattern of white tubercles; no banding on base of thigh; 9–11 dark caudal bands on original tail; white caudal bands generally not immaculate; hatchlings and juveniles bearing white tail tips; and adult posterior caudal region white. All these characters are scored across all species of the C. pulchellus complex in Table 5 in Wood et al. 2020.

Comparisons. Cyrtodactylus evanquahi sp. nov. differs from all other species in the C. pulchellus complex by having prominent tuberculation, more dark body bands, and a smaller maximum SVL (Table 5). It is further differentiated from all other species by having a combination of prominent tuberculation on the body; no tubercles on the ventral surface of the forelimbs, gular region, or in the ventrolateral folds; 31–34 paravetebral dorsal tubercles; 18–23 longitudinal rows of tubercles; 29–33 ventral scales; 22–23 subdigital lamellae on the fourth toe; 32–36 femoroprecloacl pores; a shallow precloacal groove in males; body bands and nuchal loop edged with a thin white line bearing tubercles; no scattered white sports on the dorsum; six or seven dark body bands thinner than interspaces; 9–11 dark caudal bands on original tail; bands on the original tail separated by not imperfect white caudal bands (Table 4). Additional comparisons between C. evanquahi sp. nov. and other members of the C. pulchellus complex can be found in Table 4.
Within the C. pulchellus complex, C. evanquahi sp. nov. is the sister species to C. pulchellus and can be further differentiated from it by having prominent tuberculation as opposed to moderate–moderate+; shallow precloacal groove in males opposed to a deep groove; having more dark dorsal body bands (six or seven vs. four); having thinner dark body bands; hatchlings and juveniles with white tail tips as opposed to no white tail tips; white caudal bands in adults not immaculate; adults having a posterior caudal region white (Table 5). 
EtymologyThe specific epithet honors Dr. Evan S. H. Quah, who suggested the urgency to document the herpetofaunal diversity of Gunung Baling, Kedah, which is under constant threat from two cement companies that are actively mining the limestone. He also participated in the only herpetological survey of the area. Dr. Quah is an extremely productive contributor to the study of herpetology in Malaysia and is a champion for conservation of the region. 
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Termprayoon, K., Rujirawan, A., Grismer, L. L., Wood Jr, P. L., & Aowphol, A. 2021. Taxonomic reassessment and phylogenetic placement of Cyrtodactylus phuketensis (Reptilia, Gekkonidae) based on morphological and molecular evidence. ZooKeys, 1040, 91 - get paper here
  • WOOD JR, PERRY L.; L. LEE GRISMER, MOHD ABDUL MUIN, SHAHRUL ANUAR, JAMIE R. OAKS 2020. A new potentially endangered limestone-associated Bent-toed Gecko of the Cyrtodactylus pulchellus (Squamata: Gekkonidae) complex from northern Peninsular Malaysia. Zootaxa 4751 (3): 437–460 - get paper here
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