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Cyrtodactylus kohrongensis GRISMER, ONN, OAKS, NEANG, SOKUN, MURDOCH, STUART & GRISMER, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Koh Rong Island Bent-toed Gecko 
SynonymCyrtodactylus kohrongensis GRISMER, ONN, OAKS, NEANG, SOKUN, MURDOCH, STUART & GRISMER 2020 
DistributionCambodia (Koh Rong Island, Preah Sihanouk Province)

Type locality: Monkey Beach, Koh Rong Island, Preah Sihanouk Province, Cambodia (10.69071°W 103.29003°N WGS at 16 m in elevation) .  
Reproduction 
TypesHolotype. LSUHC 10564, Adult male, collected by Jamie R. Oaks, Chan Kin Onn, Jesse L. Grismer, L Lee Grismer and Lang Sokun on 25 March 2012.
Paratypes. Paratypes were collected by the same researchers as the holotype. Juvenile female LSUHC 10545 and adult females LSUHC 10536 and CBC 03194 bear the same collection data as the holotype except that they were collected on 22 March 2012. Subadult female NCSM 98718 bears the same collection data as the holotype except collected at 10.69071°N 103.29003°E, 31 m elevation, on 21 March 2012. Adult females LSUHC 10562–63 and CBC 04189 and juvenile LSUHC 10565 were collected on Sok San Beach, Koh Rong Island, Koh Kong Province, Cambodia (10.7079° W 103.2288°N WGS at 45 m in elevation) on 25 March 2012. 
DiagnosisDiagnosis. Cyrtodactylus kohrongensis sp. nov. can be separated from all other species of the C. intermedius group by having 8–10 supralabials, 7–10 infralabials, 25–33 paravertebral tubercles, 16–20 rows of longitudinally arranged tubercles, 38–42 ventrals, 7–9 expanded subdigital lamellae on the fourth toe, 12–15 unexpanded subdigital lamellae on the fourth toe, 20–24 total subdigital lamellae on the fourth toe, 14–26 enlarged femorals, 7–9 enlarged precloacals, two or three rows of enlarged post-precloacals, two postcloacal tubercles in males, seven precloacal pores in males, enlarged femorals and enlarged precloacals not continuous, proximal femorals less than one-half the size of the distal femorals, no digital pockets, maximum SVL 76.1 mm, no large dark blotches on top of head, four well-defined brown body bands edged with white tubercles, and body bands narrower than the beige interspaces (Tables 5, 6, Grismer et al. 2020).

Comparisons. Based on meristic data, Cyrtodactylus kohrongensis sp. nov. can be differentiated from all other member of the C. intermedius group by having a significantly greater mean number of expanded lamellae on the fourth toe (8.4 versus 5.2–6.5, collectively; Fig. 3, Tables 2, 6) and a greater mean number of total fourth toe lamellae (21.9 versus 16.3–20.0, respectively; Fig. 3, Tables 2, 6). It differs further from C. auralensis, C. bokorensis, C. cardamomensis, C. phuquocensis, and C. septimontium by having a significantly lower mean number of enlarged femoral scales (19.4 versus 24.6–27.9, collectively; Fig. 3, Tables 2, 6). From its closest relative C. cardamomensis, it differs in having a significantly greater mean number of supralabials (8.9 versus 7.6; Fig. 3, Tables 2, 6) and significantly lower mean number of infralabials (8.0 versus 9.5; Fig. 3, Tables 2, 6) and precloacal scales (8.0 versus 9.5; Fig. 3, Tables 2, 6). Additional differences are illustrated in Figure 3.
Based on the scaled mensural data, Cyrtodactylus kohrongensis sp, nov. can be differentiated from all other members of the C. intermedius group by having significantly a narrower interorbital distance (Fig. 5, Table 3) and a significantly greater axilla-groin length (Fig. 5, Table 3). From its closest relative C. cardamomensis, it differs significantly in having a wider and longer head, a larger eye, and longer limbs (Fig. 5, Table 3). Additional differences are illustrated in Figure 3.
Based on color pattern and morphological data other than meristics and mensurals, Cyrtodactylus kohrongensis sp. nov. can be differentiated from all other species except C. laangensis by lacking as opposed to having continuous enlarged femoral and precloacal scales. From C. auralensis, C. bokorensis, C. laangensis, C. phuquocensis, and C. septimontium it can be differentiated by its proximal femoral scales being the same size as its distal femoral scales as opposed to being less than one-half their size. Cyrtodactylus kohrongensis sp. nov. differs further from C. auralensis by lacking as opposed to having dark pigmented blotches on the top of the head (Grismer et al. 2020). 
Comment 
EtymologyThe specific epithet kohrongensis is a Latinized toponymic adjective named after Koh Rong Island, Preah Sihanouk Province, Vietnam [sic!]. 
References
  • GRISMER, L. LEE; CHAN KIN ONN, JAMIE R. OAKS, THY NEANG, LANG SOKUN,<br />MATTHEW L. MURDOCH, BRYAN L. STUART & JESSE L. GRISMER 2020. A new insular species of the Cyrtodactylus intermedius (Squamata: Gekkonidae) group from Cambodia with a discussion of habitat preference and ecomorphology. Zootaxa 4830 (1): 075–102 - get paper here
 
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