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Cyrtodactylus siahaensis PURKAYASTHA, LALREMSANGA, LITHO, RATHEE, BOHRA, MATHIPI, BIAKZUALA & MUANSANGA, 2022

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus siahaensis PURKAYASTHA, LALREMSANGA, LITHO, RATHEE, BOHRA, MATHIPI, BIAKZUALA & MUANSANGA 2022 
DistributionIndia (Mizoram)

Type locality: Mizoram state, Siaha district, Siaha town, New Colony-I, opposite to residential buildings nearby Meisavaih road; 22.490565°N, 92.9789525°E; 1025 m a.s.l.; 13 Jul. 2021  
Reproduction 
TypesHolotype: MZMU2445, adult male, 13 Jul. 2021; 
DiagnosisDiagnosis: Cyrtodactylus siahaensis sp. nov. is a moderate-sized gecko (adult SVL 42.6–63.9 mm); 9–11 supralabials;9–11 infralabials; dorsal tubercles are rounded, bluntly conical and feebly keeled in 22–24 longitudinal rows; 36–39 paravertebral tubercles between the level of the axilla and the level of the groin; 34–37 mid-ventral scale rows; 7 precloacal pores in males; 15–18 subdigital lamellae under IV toe; no single row of transversely enlarged subcaudal scales; dorsal markings are dark brown, irregular blotches; tail with alternating dark and light bands.

Comparisons: Genetically, Cyrtodactylus siahaensis sp. nov. is a sister to C. bengkhuaiai differing from each other by an uncorrected p-distance of 8.33% as in Fig. 2 and Table 2. Morphological and meristic measurements are as shown in Table 3. Cyrtodactylus siahaensis sp. nov. has been compared morphologically with all known members of the genus Cyrtodactylus within the khasiensis group. Morphologically, Cyrtodactylus siahaensis sp. nov. can be differentiated from C. exercitus sp. nov. by having a larger maximum adult size, 72.0 mm max. (vs 68.0 mm max. in Cyrtodactylus exercitus sp. nov.), seven precloacal pores in a
continuous series vs 11–15 precloacal pores in C. exercitus and 36–39 paravertebral tubercles between the level of axilla and groin vs 32–34 tubercles in C. exercitus. Cyrtodactylus siahaensis sp. nov. differs from other members of the C. khasiensis group by the following characters: a moderate body size with a maximum SVL of 72.0 mm max. vs 81.7 mm in C. arunachalensis, 78 mm. in C. ayeyarwadyensis, 81.6 mm in C. aunglini, 77.0 mm in C. bapme, 88.0 mm in C. brevidactylus, 83.8 mm in C. chrysopylos, 62.3 mm in C. gansi, 64.5 mm in C. himalayicus, 96.2 mm in C. jaintiaensis, 80 mm. in C. kazirangaensis, 81.1 mm in C. khasiensis, 61.7 mm in C. mandalayensis, 82 mm in C. martinstolli, 78.2 mm in C. montanus, 59.0 mm in C. myaleiktaung and 90 mm in C. tamaiensis; 7 PcP in a continuous series in males vs 10–28 PcP in C. ayeyarwadyensis, 11–18 PcP in C. agarwali, 12–13 PcP in C. aunglini, 5–7 PcP in C. bengkhuaiai, 8 or 9 PcP in C. brevidactylus, 8–13 PcP in C. chrysopylos, 10 in C. himalayicus, 11–12 PcP in C. jaintiaensis, 10–11 PcP in C. kazirangaensis, 10–12 PcP in C. khasiensis, 0–8 PcP in C.martinstolli, 9–11 PcP in C. mombergi, 8–10 PcP in C. montanus, 14 PcP in C. septentrionalis, 9–12 PcP in C. urbanus, 16–29 PcFP in C. gansi, 26–39 PcFP in C. guwahatiensis, 34–38 PcFP in C. karsticola, 40 PcFP in C. tamaiensis, 29–37 PcFP in C. tripuraensis, ; 22–24 rows of tubercles throughout the dorsum vs 24–26 in C. arunachalensis, 27–30 rows in C. brevidactylus, 16–20 rows in C.chrysopylos, 19–21 rows in C. himalayicus, 19–20 rows in C. jaintiaensis, 18 rows in
C. cayuensis and C. mandalayensis, 14–15 rows in C. markuscombaii, 16–18 rows in C. nagalandensis, 21 rows in C. tamaiensis, 19–21 rows in C. tripuraensis, 34–37 rows of scales across the venter vs 38 rows in
C. arunachalensis, 47–49 rows in C. aunglini, 37–42 rows in C. bengkhuaiai, 28–34 rows in C. cayuensis, 39–55 rows in C. chrysopylos, 40–42 rows in C. jaintiaensis, 37–43 rows in C. kazirangaensis, 32 rows in C. mandalayensis, 38–39 rows in C. markuscombaii, 57 rows in C. myaleiktaung and 30–34 in C. urbanus; 36–39 paravertebral tubercles between the level of axilla and groin vs 30–35 in C. chrysopylos and 30–34 in C. jaintiaensis; 15–18 subdigital lamellae beneath toe IV vs 19–22 subdigital lamellae in C. khasiensis and C. mombergi; 13 in C. martinstolii and 10 in C. gansi; 8–10 dark blotches in the dorsum vs 11–12 blotches in C. aaronbaueri, six in C. myaleiktaung; indistinct light and dark blotches in C. karsticola and C. nagalandensis and an indistinct pattern of thick dark reticulations enclosing lighter
blotches in C.montanus; a higher TRL/SVL ratio (min. 0.47 max. 0.52 avg.0.5 vs min. 0.42 max. 0.48 avg. 0.45 in C.agarwali, min.0.35 max. 0.46 avg.0.43 in C.guwahatiensis, min. 0.44 max. 0.47 avg. 0.46 in C.septentrionalis, min 0.43, max 0.49, avg 0.46 in C.bengkhuaiai, min. 0.43 max. 0.47 avg. 0.45 in C.bapme). Cyrtodactylus siahaensis sp. nov. can again be differentiated from C. aaronbaueri by having a lower TRL/SVL ratio, min. 0.47 max. 0.52 avg.0.5 vs min 0.5, max 0.56, avg 0.52 in C. aaronbaueri. Cyrtodactylus siahaensis sp. nov. also differs from C.bapme by having 0–3 pitted scales in females (vs 0–13 precloacal pits in the latter). Furthermore, Cyrtodactylus siahaensis sp. nov. can again be differentiated from C. ayeyarwadyensis by the absence of white punctuations bordering the dark dorsal
spots in the dorsum (vs presence of dark blotches usually bordered posteriorly by white punctuations in C. ayeyarwadyensis). In Cyrtodactylus siahaensis sp. nov. the underneath of the tail with small uniformscale vs enlarged median scale line in C. cayuensis, C. khasiensis, C. martinstolli. 
Comment 
EtymologyNamed after the name of the town, Siaha (a district capital of Siaha District) from where the type series was collected. 
References
  • Purkayastha, J., Lalremsanga, H. T., Litho, B., Rathee, Y. S., Bohra, S. C., Mathipi, V., ... & Muansanga, L. 2022. Two new Cyrtodactylus (Squamata, Gekkonidae) from Northeast India. European Journal of Taxonomy, 794: 111-139
 
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