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Cyrtodactylus ywanganensis GRISMER, WOOD, THURA, QUAH, M GRISMER, MURDOCH, ESPINOZA & LIN, 2018

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common Names 
SynonymCyrtodactylus ywanganensis GRISMER, WOOD, THURA, QUAH, M GRISMER, MURDOCH, ESPINOZA & LIN 2018 
DistributionMyanmar (Shan)

Type locality: 2.7 km southwest of Ywangan, Ywangan Township, Taunggyi District, Shan State, Myanmar (21.14643°N, 96.42178°E; 1157 m in elevation)  
TypesHolotype. LSUHC 13758, adult female collected on 24 October 2017 at 2300 hrs by L. Lee Grismer, Robert E. Espinoza, Matthew L. Murdoch, Myint Kyaw Thura, Evan S. H. Quah, and Tun Oo.
Paratypes. Adult female paratype LSUHC 13757 bears the same collection data as the holotype. Adult male paratype LSUHC 13712 and subadult male paratype LSUHC 13711 bear the same collection data as the holotype except they were collected on 23 October 2017 at 2200 hrs. 
DiagnosisDiagnosis. Cyrtodactylus ywanganensis sp. nov. differs from all congeners by having the unique combination of a maximum SVL of 96.1 mm; seven supralabials; six infralabials; 32–35 paravertebral tubercles; 21–25 longitudinal rows of body tubercles; 34–36 ventral scales; relatively long digits with seven or eight expanded subdigital lamellae proximal to the digital inflection on the fourth toe, 12–14 unmodified distal subdigital lamellae, 19–21 total subdigital lamellae; raised, strongly keeled, dorsal body tubercles; tubercles not extending beyond base of tail; enlarged femoral and precloacal scales continuous; enlarged proximal femoral scales equal in size to the enlarged distal femoral scales; 28–30 enlarged femoral scales; 20 femoral pores in male; eight or nine precloacal pores in male; femoral and precloacal pore-bearing scales not continuous; eight or nine enlarged precloacal scales; three rows of enlarged post-precloacal scales; medial subcaudal scales twice as wide as long, not extending onto lateral surface of tail; top of head darkly blotched in adults, no yellow reticulum; nuchal loop not divided medially, pronounced anterior azygous notch, posterior border straight; four jagged, dark, dorsal bands lacking paravertebral elements, wider than interspaces with no lightened centers, edged with yellow tubercles; no band on nape; dark markings in dorsal interspaces; ventrolateral folds not whitish; anterodrosal margins of thighs and brachia darkly pigmented; seven light caudal bands bearing dark markings, encircling tail; seven dark caudal bands wider than light caudal bands; and mature regenerated tail spotted. These characters are scored against all other species in the linnwayensis group (Table 3) and all other Burmese species of the Indochinese clade in Grismer et al. 2017:Table 8.

Comparisons. The phylogenetic relationships indicate that Cyrtodactylus ywanganensis sp. nov. is nested within the linnwayensis group. The PCA and DAPC analyses indicate that C. ywanganensis sp. nov. is morphospatially separate from other species of the linnwayensis group along the first three principal components (PC) with PC1 accounting for 41% of the total variation in the data set (Fig. 4; Table 5) and loading most heavily for numbers of paravertebral tubercles (PVT), expanded and total numbers of fourth toe lamellae (TL and TEL), longitudinal rows of tubercles (LRT), body bands (BB), and post-precloacal scale rows (PPS). PC2, which accounts for 18% of the total variation loads most heavily for infralabials (IL) and supralabials (SL). PC 3 accounts for an additional 11% of the total variation and loads most heavily for the number of unmodified toe lamellae (TLN). The ANOVA and Tukey HSD analyses indicate C. ywanganensis sp. nov. has several meristic characters whose mean values differ significantly (p ≤ 0.05) and discreetly (i.e. no overlap in the range of variation) from the other members of the linnwayensis group (Tables 3, 6).
EtymologyThe specific epithet, ywanganensis, is a noun in apposition in reference to the type locality being near the town of Ywangan, Shan State. 
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD JR., EVAN S. H. QUAH, MYINT KYAW THURA,<br />JAMIE R. OAKS & AUNG LIN 2019. A new species of Bent-toed Gecko (Squamata, Gekkonidae, Cyrtodactylus) from the Shan Plateau in eastern Myanmar (Burma). Zootaxa 4624 (3): 301–321 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, JR., MYINT KYAW THURA, EVAN S. H. QUAH,<br />MARTA S. GRISMER, MATTHEW L. MURDOCH, ROBERT E. ESPINOZA & AUNG LIN 2018. A new Cyrtodactylus Gray, 1827 (Squamata, Gekkonidae) from the Shan Hills and the biogeography of Bent-toed Geckos from eastern Myanmar. Zootaxa 4446 (4): 477–500 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR., MYINT KYAW THURA, THAW ZIN, EVAN S. H. QUAH, MATTHEW L. MURDOCH, MARTA S. GRISMER, AUNG LIN, HTET KYAW and NGWE LWIN 2017. Twelve new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from isolated limestone habitats in east- central and southern Myanmar demonstrate high localized diversity and unprecedented microendemism. Zoological Journal of the Linnean Society 182: 862-959 - get paper here
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