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Dendragama boulengeri DORIA, 1888

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Boulenger's Tree Agama 
SynonymDendragama boulengeri DORIA 1888
Dendragama boulengeri — WERNER 1900: 484
Dendragama boulengeri — DE ROOIJ 1915: 118
Calotes (Pseudocalotes) boulengeri — MERTENS 1954: 185
Calotes boulengeri — WERMUTH 1967: 34
Calotes boulengeri — BRYGOO 1988: 9
Dendragama boulengeri — MANTHEY & GROSSMANN 1997: 167
Dendragama boulengeri — MANTHEY & SCHUSTER 1999: 51
Dendragama boulengeri — TEYNIÉ et al. 2010
Dendragama schneideri — HARVEY Et al. 2017 
DistributionIndonesia (Sumatra)

Type locality: Mount Singalang, West Sumatra (elevation 2800 feet) = Gunung Singgalang, Kabupaten Agam, Sumatera Barat, Indonesia, 0.38°S, 100.36°E  
TypesLectotype: MSNG 29936, male; paralectotype (?): MRSN (= MSNTO) R2856; MNHN-RA MHNP 1889.27 (2133 alpha) fide Brygoo 1988. 
DiagnosisDiagnosis: The following combination of characters distinguishes Dendragama boulengeri from its congeners: (1) enlarged scales below eye separated from supralabials by continuous row of small lorilabials; (2) enlarged tympanic scale intermediate in size (7–14% of head length) and set far back on head (orbit–meatus 24–32% of head length); (3) sublabial tubercular scales 0–3, usually 1, in front of postrictal modified scale; (4) gulars small, 30–38; (5) dorsal crest of 11–18 projecting scales, serrate with obvious gaps between enlarged projecting scales; (6) dorsolateral crest of more or less continuous posttympanic and dorsolateral series; (7) scales around midbody 62–84; (8) scales of lower flanks heterogenous in size, smooth to feebly keeled; (9) ventrals heavily keeled, 50–63; (10) no white sublabial stripe extending from below eye to level of ear, however 1–3 white spots usually present in this region; (11) most specimens with prominent black prescapular blotch edged posteriorly in white; (12) proximal half of tail with 7–9 dark brown or green bands; (13) buccal epithelium and tongue yellow or orange [Harvey et al. 2017: 83] 
CommentSynonymy partly after WERMUTH 1967. Shaney et al. 2020 formally revalidated D. schneideri from the synonymy of D. boulengeri.

Type species: Calotes boulengeri DORIA 1888 is the type species of the genus Dendragama DORIA 1888.

Key: Shaney et al. 2020 provide a key to the species of Dendragama.

Distribution: see map in Shaney et al 2020: 130 (Fig. 1). 
EtymologyNamed after George Albert Boulenger (1858-1937), herpetologist at the British Museum of Natural History, London. 
  • Ahl, E. 1926. Neue Eidechsen und Amphibien. Zool. Anz. 67: 186-192
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Brygoo, E.R. 1988. Les types d’Agamidés (Reptiles, Sauriens) du Muséum national d’Histoire naturelle Catalogue critique. Bull. Mus. Nat. Hist. Nat. 10 (ser. 4) A (3), suppl.: 1-56
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Doria, G. 1888. Note Erpetologiche - Alcuni nuovi Sauri raccolti in Sumatra dal Dr. O. Beccari. Ann. Mus. civ. stor. nat. Genova 2 (ser. 6): 646-652 - get paper here
  • Hallermann, J. 2005. The bizarre arboreal agamids. Reptilia (GB) (42): 8-15 - get paper here
  • Hallermann, J. 2005. Mit Hörnern, Kämmen und Gleithäuten - die bizarren Baumagamen. Reptilia (Münster) 10 (51): 18-25 - get paper here
  • Harvey, Michael B.; Kyle Shaney, Irvan Sidik, Nia Kurniawan, and Eric N. Smith 2017. Endemic Dragons of Sumatra’s Volcanoes: New Species of Dendragama (Squamata: Agamidae) and Status of Salea rosaceum Thominot. Herpetological Monographs 31 (1): 69–97 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • Mertens, R. 1954. Über die javanische Eidechse Dendragama fruhstorferi und die Gattung Dendragama. Senckenbergiana 34 (4/6): 185-186
  • Prötzel, D. & J. Forster 2014. Herpetologische Eindrücke von Sumatra. Reptilia (Münster) 19 (109): 80-85
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Werner, F. 1900. Reptilien und Batrachier aus Sumatra, gesammelt Herrn. Gustav Schneider jr., im Jahre 1897-1898. Zool. Jahrb. (syst) 13: 479-508 - get paper here
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