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Dendrelaphis levitoni VAN ROOIJEN & VOGEL, 2012

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Higher TaxaColubridae, Ahaetuliinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Leviton’s Bronze-back Tree Snake 
SynonymDendrelaphis levitoni VAN ROOIJEN & VOGEL 2012
Dendrelaphis caudolineatus — BOULENGER 1894: 90
Dendrelaphis caudolineatus — GRIFFIN 1909: 598
Dendrelaphis caudolineatus — GRIFFIN 1911: 261
Dendrelaphis caudolineatus — TAYLOR 1922: 172
Dendrelaphis caudolineatus caudolineatus — MEISE & HENNIG 1932: 282
Dendrelaphis caudolineatus caudolineatus — LEVITON 1963: 378, 401
Dendrelaphis caudolineatus caudolineatus — LEVITON 1968: 382–385
Dendrelaphis levitoni — VAN ROOIJEN & VOGEL 2012
Dendrelaphis levitoni — WALLACH et al. 2014: 218 
DistributionPhilippines (Palawan, Balabac and Candaraman)

Type locality: Puerto Princesa, Palawan, P  
TypesHolotype: CAS 15803, adult female; collector A. Seale; collected 6 July 1908; Paratypes. ZMA.RENA 21570, Philippines (Palawan); BMNH 1894.6.30.48, Palawan, collected by Everett; SMF 74464, Bataraza, collected by M. Gaulke; USNM 158273, Initiam, collected by Thomson; USNM 158276, Puerto Princesa, collected by Kuntz. 
DiagnosisDescription. A species of the Dendrelaphis caudolineatus group, characterized by the combination of: 1) black longitudinal stripes along the entire length of the body, 8 stripes at midbody; 2) a pale ventrolateral stripe along the entire length of the body; 3) a black postocular stripe which extends onto the neck where it becomes one the upper longitudinal stripe that borders the pale ventrolateral line; 4) 175–181 ventrals (males), 180–189 ventrals (females); 5) 101–116 divided subcaudals; 6) relative tail-length 0.250–0.285; 7) 4–6 temporal scales on each side; 8) 9–11 infralabials on each side; 9) 4–5 scales touching the posterior borders of the parietal scales; 10) maximum total length males 81.5 cm, females 130.5 cm [VAN ROOIJEN & VOGEL 2012].

Sexual dimorphism. On average, males have 177 ventral scales whereas females have 184 ventral scales (F = 5.4; df = 1; p = 0.05). Females grow larger than males [VAN ROOIJEN & VOGEL 2012]. 
CommentHabitat: fully arboreal (Harrington et al. 2018). 
EtymologyThis species is named after A. E. Leviton in honour of his extensive work on the Philippine snakes. 
  • Blackburn, D.G. 1993. STANDARDIZED CRITERIA FOR THE RECOGNITION OF REPRODUCTIVE MODES IN SQUAMATE REPTILES. Herpetologica 49 (1): 118-132 - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Kwet, Axel 2013. Liste der im Jahr 2012 neu beschriebenen Reptilien. Terraria-Elaphe 2013 (3): 52-67 - get paper here
  • ROOIJEN, JOHAN VAN & GERNOT VOGEL 2012. A revision of the taxonomy of Dendrelaphis caudolineatus (Gray, 1834) (Serpentes: Colubridae). Zootaxa 3272: 1–25 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Weinell, Jeffrey L.; Errol Hooper, Alan E. Leviton, Rafe M. Brown 2019. Illustrated Key to the Snakes of the Philippines. Proc. Cal. Acad. Sci. (4) 66 (1): 1-49 - get paper here
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