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Dipsadoboa kageleri (UTHMÖLLER, 1939)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Kageler’s Tree Snake 
SynonymDipsadoboa hotamboeia kageleri UTHMÖLLER 1939
Crotaphopeltis hotamboiea tornieri UTHMÖLLER 1937
Dipsadoboa shrevei kageleri — BROADLEY & HOWELL 1991: 31
Dipsadoboa kageleri — BRANCH et al. 2019 
DistributionTanzania (Vicinity of Mount Kilimanjaro)

Type locality: Sanya, Tanzania, 1350 m elevation.  
Reproductionoviparous 
TypesHolotype: ZSM 254/1937 (originally private collection W. Uthmöller Nr. 93), adult male 
DiagnosisDiagnosis (kageleri): A dry savanna form of Dipsadoboa which is apparently endemic to the Kilimanjaro area and which has the following character combination: dorsals in 17-17-13 rows; ventrals 191 and 195, male and female, respectively; anal divided; subcaudals 72 + x and 76 + x, male and female, respectively; 14 + II + 1 maxillary teeth, color similar to the adult color of the nominate subspecies.

Dipsadoboa shrevei kageleri inhabits the most xeric habitat of any member of the genus, and also shows a combination of characters that differentiate it from all known Dipsadoboa, and particularly from D. s. shrevei. These include 17-17-13 dorsal scale rows (19-19-15 in D. s. shrevei), lower ventral counts in males (191 vs. 203–210 in D. shrevei) and females (195 vs. 199–210 in D. s. shrevei), lower subcaudal counts in males (72 v 74–91 in D. s. shrevei) and females (68–76 vs. 75–91 in D. s. shrevei; excluding high counts for Mtene females, see below), a divided anal (entire in D. s. shrevei), and possibly size (total length 670 mm vs. 1241 mm in D. s. shrevei). The genetic material available also shows species level divergence from all other Dipsadoboa sampled (Table 2, Fig. 2), notably that it is divergent from typical D. s. shrevei (16.5% for cyt b), and supports the considerable morphological differences between the two taxa (Table 4, Branch et al. 2019).  
CommentSynonymy: mainly after Rasmussen 1986. 
EtymologyNamed after the owner of the coffee plantation where the type was found, Mr. G. Kageler. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Berry, P. S. M. 1970. A Note on Shreve's Tree Snake Dipsadoboa shrevei in Zambia. J. Herp. Assoc. Africa (6): 20-21 - get paper here
  • BRANCH, WILLIAM R.; JULIAN BAYLISS, GABRIELA B. BITTENCOURT-SILVA, WERNER CONRADIE, HANLIE M. ENGELBRECHT, SIMON P. LOADER, MICHELE MENEGON, CRISTÓVÃO NANVONAMUQUITXO, KRYSTAL A. TOLLEY 2019. A new species of tree snake (Dipsadoboa, Serpentes: Colubridae) from ‘sky island’ forests in northern Mozambique, with notes on other members of the Dipsadoboa werneri group. Zootaxa 4646 (3): 541–563 - get paper here
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Broadley, D.G. 1958. Occurance of Dipsadoboa shrevei in Northern Rhodesia. J. Herp. Assoc. Rhodesia 2: 5 - get paper here
  • Broadley, D.G. 1991. The Herpetofauna of Northern Mwinilunga Distr., Northw. Zambia. Arnoldia Zimbabwe 9 (37): 519-538
  • Broadley, D.G. 1998. The reptilian fauna of the Democratic Republic of the Congo (Congo-Kinshasa). In: Schmidt, K.P. and Noble, G.K., Contributions to the Herpetology of the Belgian Congo... [reprint of the 1919 and 1923 papers]. SSAR Facsimile reprints in Herpetology, 780 pp.
  • Broadley, D.G.; Doria, C.T. & Wigge, J. 2003. Snakes of Zambia. An Atlas and Field Guide. Edition Chimaira, Frankfurt, 280 pp. [review in Sauria 26 (3): 21]
  • Broadley, Donald G. and F. P. D. Cotterill. 2004. The reptiles of southeast Katanga, an overlooked 'hot spot'. [Congo]. African Journal of Herpetology 53 (1): 35-61. - get paper here
  • Chifundera, K. 1990. Snakes of Zaire and their bites. Afr. Stud. Monogr. (Kyoto) 10(3): 137-157.
  • Chippaux, Jean-Philippe & Kate Jackson 2019. Snakes of Central and Western Africa. Johns Hopkins University Press, 448 pp. - get paper here
  • Haagner,G.V.; Branch,W.R. & Haagner,A.J.F. 2000. Notes on a collection of reptiles from Zambia and adjacent areas of the Democratic Republic of the Congo. Annals of the Eastern Cape Museum 1: 1 – 25
  • Loveridge, A. 1932. New opisthoglyphous snakes of the genera Crotaphopeltis and Trimerorhinus from Angola and Kenya Colony. Proc. Biol. Soc. Washington 45: 83-86 - get paper here
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • Rasmussen,J.B. 1986. On the taxonomic status of Dipsadoboa werneri (BOULENGER), D. shrevei (LOVERIDGE), and Crotaphopeltis hotamboiea kageleri UTHMÖLLER (Boiginae, Serpentes). Amphibia-Reptilia 7: 51-73 - get paper here
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Trape, J.F. & R. ROUX-ESTÈVE 1995. Les serpents du Congo: liste commentée et clé de détermination. Journal of African Zoology 109 (1): 31-50
  • Uthmöller,W. 1939. Über eine neue Rasse von Crotaphopeltis hotamboiea. Zool. Anz. 125: 108-112
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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