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Higher TaxaGymnophthalmidae (Ecpleopodinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Common NamesPortuguese: Lagartinho-do-Folhiço 
DistributionBrazil (Paraiba, Bahia, Pernambuco)

Type locality: João Pessoa (07°07’S, 34°52’W): state of Paraiba: Brazil  
TypesHolotype: MZUSP 60635, an adult male, collected by Miguel T. Rodrigues 6 September 1983. 
DiagnosisDefinition (genus Dryadosaura): Small gymnophthalmid with a robust and short head, elongate body and short, stout, pentadactyl limbs. Collar fold, ear opening and eyelid present. Frontonasal single; prefrontals, parietals and interparietals present; frontoparietals absent. Two pairs of chin shields and three supraoculars. Dorsal scales in transverse rows; anteriorly smooth, imbricate, rounded in the occipital region, becoming narrower, elongate and rectangular at the arm level; posterior to the arm, dorsal scales become progressively and irregularly mucronate, with lateral sides almost juxta- posed, keeled. Lateral scales of body about the same size as dorsals, smooth, rectangular, and laterally imbricate. Ventral scales smooth, juxtaposed or slightly imbricate, rectangular. Males with preanal and femoral pores; only preanal pores in females.

For diagnostic morphological characters distinguishing Marinussaurus, Amapasaurus, Anotosaura, Arthrosaura, Colobosauroides, Dryadosaura,Ecpleopus, Kaieteurosaurus, Leposoma, and Pantepuisaurus see Table 2 in PELOSO et al. 2011.

Diagnosis (of species): Microteiid characterized by an elongate body, five toes and fingers, and short stout limbs. Collar fold, ear opening, and eyelid present. Frontonasal single; prefrontals, parietals and interparietals present; frontoparietals absent. Two pairs of chin shields; three supraoculars. Dorsal scales in 23–25 regular transversal rows; anteriorly smooth, becoming progressively hexagonal and mucronate; keeled at rump level. Lateral scales similar to dorsals, smooth, rectangular, lat- erally imbricate. Ventral scales smooth, juxtaposed or slightly imbricate, rectangular, in 15–17 rows; 26–33 scales around midbody. Infradigital lamellae mostly divided 6–9 in finger IV, and 13–16 in toe IV. Preanal and femoral pores number four and six, respectively, in males; 2-4 preanal pores in females. 
CommentThis specis and genus is most closely related to Anotosaura sp.

Type species: Dryadosaura nordestina RODRIGUES et al. 2005 is the type species of the genus Dryadosaura RODRIGUES et al. 2005. 
EtymologyFrom the Greek Dryades, meaning ‘wood-land nymph’. This was the first name proposed (by Carl Philipp von Martius) for the biogeographical province of the Atlantic Forest. “Nordestina” referes to the type locality, the north-eastern part of Brazil. 
  • ANDRADE LIMA, J.H., DIAS, E.G., COSTA, R.D.L., SILVA, F.J., LIMA, E.S.M., SANTOS, E.M., KOKUBUM, M.N.C. 2021. Lizards and snakes of Refúgio de Vida Silvestre Matas do Siriji, an Atlantic Forest hotspot of the Pernambuco Endemism Center, Northeastern Brazil. Biota Neotropica 21 (2): e20201106 - get paper here
  • Bezerra, P. E. S., Santana, F. M., & de Moura, G. J. B. 2021. Sustainability assessment of a lizard assemblage in Pernambuco state, Brazil. Journal for Nature Conservation, 125957
  • Dubeux, M. J. M., Araújo Neto, J. V. D., Triburcio, I. C. S., Lisboa, B. S., Torquato, S., Freitas, M. A. D., ... & Mott, T. 2022. A “hotspot” within a hotspot: the reptiles of the Estação Ecológica and Área de Proteção Ambiental de Murici, Atlantic Forest of northeastern Brazil. Biota Neotropica, 22: 1-14 - get paper here
  • Franzini, Lissa Dellefrate; Izabel Regina Soares Silva, Daniel Oliveira Santana, Fagner Ribeiro Delfim, Gustavo Henrique Calazans Vieira, Daniel Oliveira Mesquita 2019. Lizards fauna from the state of Paraíba, Northeastern Brazil: current knowledge and sampling discontinuities. Herpetology Notes 12: 749-763 - get paper here
  • Freitas, Marco Antonio de 2014. Squamate reptiles of the Atlantic Forest of northern Bahia, Brazil. Check List 10 (5): 1020-1030 - get paper here
  • Gonzalez R. C. et al. 2020. Lista dos Nomes Populares dos Répteis no Brasil – Primeira Versão. Herpetologia Brasileira 9 (2): 121 – 214 - get paper here
  • Melo, Íris Virgínea; Geraldo Jorge Barbosa de Moura, Marco Antonio de Freitas, Edson Victor Euclides de Andrade, Cláudio Casal, Arthur Diesel Abegg, Marcelo Nogueira de Carvalho Kakubum 2018. New additions to herpetofauna of the Parque Estadual Dois Irmãos, an urban Atlantic Rainforest Fragment, Recife municipality, Pernambuco state, northeastern Brazil. Herpetology Notes 11: 245-254 - get paper here
  • Mesquita, Daniel Oliveira; Brygida Alves, Carmem Pedro, Daniel Laranjeiras, Francis Caldas, Isabella Pedrosa, Jefter Rodrigues, Leandro Drummond, Lucas Cavalcanti, Milena Wachlevski, Paulo Nogueira-Costa, Rafaela França, Frederico França 2018. Herpetofauna in two habitat types (tabuleiros and Stational Semidecidual Forest) in the Reserva Biológica Guaribas, northeastern Brazil. Herpetology Notes 11: 455-474 - get paper here
  • Nascimento de Oliveira, Camila; Sérgio Luiz da Silva Muniz, Geraldo Jorge Barbosa de Moura 2016. Reptiles of an urban Atlantic Rainforest fragment in the state of Pernambuco, northeastern Brazil. Herpetology Notes 9: 175-183 - get paper here
  • OLIVEIRA, P.M.A., MELLO, A.V.A., DUBEUX, M.J.M., OLIVEIRA, S.B.A., LOURENÇO, G.F., NUNES, P.M.S. 2021. Herpetofauna of Matas de Água Azul, an Atlantic Forest remnant in Serra do Mascarenhas, Pernambuco state, Brazil. Biota Neotropica 21 (2): e20201063 - get paper here
  • Peloso, Pedro L.V.; Katia C. M. Pellegrino, Miguel T. Rodrigues, and Teresa C.S. Ávila-Pires 2011. Description and Phylogenetic Relationships of a New Genus and Species of Lizard (Squamata, Gymnophthalmidae) from the Amazonian Rainforest of Northern Brazil. American Museum Novitates (3713): 1-24 - get paper here
  • Roberto, I. J., R. W. Ávila & A. R. Melgarejo 2015. Répteis (Testudines, Squamata, Crocodylia) da reserva biológica de Pedra Talhada. In : studer, a., l. nusbaumer & r. spichiger (eds.). biodiversidade da reserva biológica de Pedra talhada (alagoas, Pernambuco - brasil). Boissiera 68: 357-375 - get paper here
  • Roberto, Igor Joventino; Cícero Ricardo de Oliveira, João Antonio de Araújo Filho, Herivelto Faustino de Oliveira, Robson Waldemar Ávila 2017. The herpetofauna of the Serra do Urubu mountain range: a key biodiversity area for conservation in the brazilian atlantic forest. Papéis Avulsos de Zoologia 57 (27): 347-373 - get paper here
  • RODRIGUES, MIGUEL TREFAUT; ELIZA MARIA XAVIER FREIRE, KATIA CRISTINA MACHADO PELLEGRINO and JACK W. SITES JR 2005. Phylogenetic relationships of a new genus and species of microteiid lizard from the Atlantic forest of north-eastern Brazil (Squamata, Gymnophthalmidae). Zoological Journal of the Linnean Society 144 (4): 543-557 - get paper here
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