You are here » home advanced search search results Emys orbicularis

Emys orbicularis (LINNAEUS, 1758)

IUCN Red List - Emys orbicularis - Least Concern, LR

Can you confirm these amateur observations of Emys orbicularis?

Add your own observation of
Emys orbicularis »

Find more photos by Google images search: Google images

Higher TaxaEmydidae, Emydinae, Testudinoidea, Testudines (turtles)
SubspeciesEmys orbicularis orbicularis (LINNAEUS 1758)
Emys orbicularis eiselti FRITZ, BARAN, BUDAK & AMTHAUER 1998
Emys orbicularis galloitalica FRITZ 1995
Emys orbicularis hellenica (VALENCIENNES 1832)
Emys orbicularis ingauna JESU, PIOMBO, SALVIDIO, LAMAGNI, ORTALE & GENTA 2004
Emys orbicularis occidentalis FRITZ 1993
Emys orbicularis persica EICHWALD 1831 
Common NamesE: European Pond Turtle
eiselti: Eiselt’s Pond Turtle
fritzjuergenobsti: Iberian Pond Turtle, Obst’s Pond Turtle
galloitalica: Franco-Italian Pond Turtle
hellenica: Hellenic Pond Turtle
ingauna: Ligurian Pond Turtle
occidentalis: Magreb Pond Turtle
persica: Eastern Pond Turtle
G: Europäische Sumpfschildkröte
NL: moerasschildpad
Croatian: barska kornjača
Russian: Болотная черепаха 
SynonymTestudo orbicularis LINNAEUS 1758: 198
Testudo lutaria LINNAEUS 1758: 198
Testudo terrestris GARSAULT 1764
Testudo europaea SCHNEIDER 1783
Testudo punctata GOTTWALD 1792 (nomen nudum)
Testudo pulchella SCHOEPFF 180x
Emys lutaria — LINK 1807: 52
Testudo rotunda MERREM 1820
Testudo lutraria GRAY 1831 (ex errore)
Cistudo hellenica VALENCIENNES 1832
Emys iberica VALENCIENNES 1832 (nomen nudum)
Emys antiquorum VALENCIENNES 1833
Emys turfa MEYER 1835:67
Cistudo Europaea — DUMÉRIL & BIBRON 1835: 220
Emys hofmanni FITZINGER 1836
Emys lutaria — BELL 1836-1842
Emys lutaria borealis NILSSON 1841:208
Emys europaea — RATHKE 1846: 316
Emys lutaria — CALORI 1853
Cistudo Europaea — DUMÉRIL & BIBRON 1854: 224
Lutremys europaea — GRAY 1856: 264
Cistudo anhaltina GIEBEL 1866:1
Emys orbicularis — BLANFORD 1876
Emys europaea — LEIDY 1888: 127
Emys lutaria taurica MEHNERT 1890
Emys tigris SALVATOR 1897 (nomen nudum)
Emys europaea var. sparsa DÜRIGEN 1897
Emys europaea var. maculosa DÜRIGEN 1897
Emys europaea var. concolor DÜRIGEN 1897
Emys europaea var. punctata DÜRIGEN 1897
Emys orbicularis aralensis NIKOLSKY 1915
Emys orbicularis luteofusca FRITZ 1989
Emys orbicularis colchica FRITZ 1994
Emys orbicularis — ENGELMANN et al. 1993
Emys orbicularis — SCHLEICH, KÄSTLE & KABISCH 1996: 142
Emys orbicularis colchica — DIETRICH 2008
Emys orbicularis — SINDACO & JEREMČENKO 2008
Emys orbicularis orbicularis — SEIDEL: & ERNST 2017

Emys orbicularis eiselti FRITZ et al. 1998
Emys orbicularis eiselti — RHODIN et al. 2010
Emys orbicularis eiselti — SEIDEL & ERNST 2017

Emys orbicularis galloitalica FRITZ 1995
Emys orbicularis galloitalica FRITZ 1995: 217
Emys orbicularis capolongoi FRITZ 1995: 204
Emys orbicularis lanzai FRITZ 1995: 211
Emys orbicularis capolongoi — MUTZ et al. 1999
Emys orbicularis capolongii [sic] — OBST 2003: 16
Emys orbicularis galloitalica — RHODIN et al. 2010
Emys orbicularis galloitalica — SEIDEL & ERNST 2017

Emys orbicularis hellenica (VALENCIENNES in BIBRON & BORY 1832)
Cistuda hellenica VALENCIENNES in BIBRON and BORY 1832:61
Emys antiquorum VALENCIENNES in BORY de SAINT-VINCENT 1833:pl.9
Emys (Emys) hofmanni FITZINGER 1835:123 (nomen novum)
Emys orbicularis hoffmani SCHREIBER 1875:537
Emys orbicularis atra WERNER 1897:15
Emys europaea maculosa DÜRIGEN 1897:15
Emys orbicularis hellenica — FRITZ 1995: 224
Emys orbicularis hellenica — RUTSCHKE et al. 2005
Emys orbicularis hellenica — RHODIN et al. 2010
Emys orbicularis hellenica — SEIDEL & ERNST 2017

Emys orbicularis ingauna JESU, PIOMBO, SALVIDIO, LAMAGNI, ORTALE & GENTA 2004
Emys orbicularis ingauna — RHODIN et al. 2010
Emys orbicularis ingauna — SEIDEL & ERNST 2017

Emys orbicularis occidentalis FRITZ 1993
Emys orbicularis fritzjuergenobsti FRITZ 1993
Emys orbicularis occidentalis — FRITZ 1995: 196
Emys orbicularis fritzjuergenobsti — FRITZ 1995: 200
Emys orbicularis hispanica FRITZ, KELLER & BUDDE 1996
Emys orbicularis fritzjurgenobsti — SINDACO & JEREMČENKO 2008
Emys orbicularis fritzjurgenobsti — RHODIN et al. 2010
Emys orbicularis occidentalis — RHODIN et al. 2010
Emys orbicularis occidentalis — TTWG 2017
Emys orbicularis occidentalis — SEIDEL & ERNST 2017
Emys orbicularis occidentalis — PEREIRA et al. 2018

Emys orbicularis persica EICHWALD 1831
Emys orbicularis iberica EICHWALD 1831
Emys orbicularis kurae FRITZ 1994
Emys orbicularis orientalis FRITZ 1994
Emys orbicularis iberica — SINDACO & JEREMČENKO 2008
Emys orbicularis persica — RHODIN et al. 2010
Emys orbicularis persica — SEIDEL & ERNST 2017 
DistributionGermany, Austria, Switzerland, Poland, Hungary, Albania, Yugoslavia, Czech Republic, Slovakia, Italy, Sardinia, S France, Corsica, Spain, Balearic Islands: Mallorca, Menorca, Portugal, Greece (including Limnos, Lesbos, Corfu = Corfou, Samothraki), Turkey, Bulgaria, Romania,
Iran (Caspian Sea), Soviet Union, Latvia, Lithuania,
Morocco, Algeria, Tunisia (Schleich et al.).

capolongoi: Sardinia; Type locality: “Olbia (Sardinien)”

colchica: NE coastal Turkey

eiselti: SE Turkey, Syria

fritzjuergenobsti (obsolete): Portugal, Spain, S France, Italy (Thyrrenian coast)

galloitalica: NE Spain, S France, Italy

hellenica: Albania, Bosnia, Hercegovina, Crotia, Macedonia, Serbia, Slovenia, Montenegro; Italy, Sicily, Greece, Aegean Turkey. Type locality: Ebene von Nisi am Fluß Pamisos, Zentral-Messina, Peloponnes (fide Fritz 1995: 224)

iberica: Caspian coast from Dagestan to E Transcaucasia

ingauna: Italy

lanzai: Corsica; Type locality: “Conca-Mündung bei Fontea, unweit Santa Lucia di Porto-Vecchio (Korsika)”

luteofusca: Turkey (C Anatolia)

occidentalis: Morocco, N Algeria and N Tunisia, Portugal (Malkmus 1995)

orbicularis: Armenia, Austria, Azerbaijan, Belarus, Bosnia and
Herzegovina, Bulgaria, Croatia, Czech Republic (extirpated), France, Republic of Georgia, Germany, Hungary, Italy,Kazakhstan, Latvia, Lithuania, Moldova, Poland,Romania, Russia, Serbia, Slovakia, Slovenia, N Spain,Turkey, Turkmenistan, Ukraine.

persica: N Iran to Turkmenistan, Armenia, Azerbaijan, Republic of Georgia, Iran, Russia (Dagestan)

Introduced to the Balearic Islands (LEVER 2003).

acording to the 1994 IUCN Red List of Threatened Animals:
Albania, Algeria, Armenia, Austria,
Azerbaijan, Belarus, Belgium, Bosnia and Hercegovina,
Bulgaria, Croatia (including some Adriatic islands),
Cyprus, Czech Republic, France, Republic of Georgia, Germany, Greece, Hungary, Iran, Kazakhstan, Latvia, Liechtenstein, Lithuania, Macedonia, Monte Negro, Moldova, Monaco, Morocco, Netherlands, Poland, Portugal, Romania, C/S Russia, Serbia, Slovakia, Slovenia, Spain, Switzerland, Tunisia, Turkey, W Turkmenistan, Ukraine, Yugoslavia

Type locality: S Europe  
ReproductionEmys orbicularis has temperature-dependent sex determination (TSD, Pieau 1998, GIRONDOT et al. 1994, SARRE et al. 2004). 
TypesHolotype: unlocated fide Iverson 1992: 177
Holotype: NMW [eiselti]
Holotype: MNHN-RA 1993.5804; paratypes: MNHN-RA 1993.5801-5803, MTD 32366,33710, NMW [galloitalica]
Holotype: MNHN-RA 1943, adult male [hellenica]
Holotype: MSNG 50650, paratypes: MSNG [ingauna]
Holotype: NMW [kurae]
Paratypes: NMW [orientalis]
Holotype: SMF 59593, Paratypes: MTD 17615, ZFMK 55578-580, 3 adult males, SMF, HSC, MZUF [capolongoi]
Holotype: MNHN-RA 1961.0344; paratypes: MTD 31815 [occidentalis] 
CommentDistribution: Postglacial remains have been found in Sweden, Denmark, the Netherlands, and England. E. o. galloitalica was introduced long ago in Corsica and Sardinia and naturalized there (Vamberger et al. 2015).

Although E. orbicularis is found in many places, it is questionable whether it breeds there (e.g. in Liechtenstein). Many animals might be escaped pets or otherwise introduced. Especially distribution records from Middle and Northern Europe should be considered with caution. The 1994 IUCN Red List of Threatened Animals lists E. orbicularis erroneously from Cyprus. E. o. galloitalica was introduced long ago in Corsica and Sardinia and naturalized there.

Subspecies: In a recent list by OBST (2003) the subspecies Emys orbicularis kurae FRITZ 1994 and E. o. orientalis FRITZ 1994 are not included. Their status remains unclear. Velo-Anton et al. (2008) synonymized hispanica with fritzjuergenobsti. Pedall et al. (2010) and Rhodin et al. (2010) synonymized capolongoi and lanzai with galloitalica.

Synonymy mainly after Kwet (2010) in Reptilia 15 (85): 86 and Rhodin et al. (2010), TTWG 2017.

Type species: Testudo europaea SCHNEIDER 1783 is the type species of the genus Emys DUMERIL 1806.

Habitat: freshwater (swamps, ponds, rivers) 
EtymologyNamed after the Greek word for freshwater turtle, “emus”. Latin “orbis” = a circle; orbiculus a little circle or disc; -aris (Latin) pertaining to. 
  • Akgül, Riza, Tugce Caprazli, Nesrin Tuurker, Hüseyin Erdugan and Murat Tosunoglu. 2014. Epizoic algae on Emys orbicularis (Linnaeus, 1758), and Mauremys rivulata (Valenciennes, 1833), in the Kavak River Delta (Saros Bay, Turkey) (Testudines: Emydidae, Geoemydidae). Herpetozoa 27 (1/2): 21-28 - get paper here
  • Akman, B., Çakmak, M., & Yıldız, M.Z. 2020. On the Herpetofauna of the Central Anatolian Province of Kırıkkale (Turkey) (Amphibia; Reptilia). Acta Biologica Turcica, 33(2): 70-79
  • Ananjeva, NB, Orlov NL; Khalikov RG; Darevsky IS; Ryabov IS & Barabanov AV 2006. The Reptiles of North Eurasia. Taxonomic Diversity, Distribution, Conservation Status [this comprises the territory of the former Soviet Union and Mongolia]. Pensoft Series Faunistica 47, 250 pp.
  • ARAKELYAN, Marine AND JAMES F. PARHAM 2008. The Geographic Distribution of Turtles in Armenia and the Nagorno-Karabakh Republic (Artsakh). Chelonian Conservation and Biology 7(1): 70–77 - get paper here
  • Arakelyan, Marine S.; Felix D. Danielyan, Claudia Corti, Roberto Sindaco, and Alan E. Leviton 2011. The Herpetofauna of Armenia and Nagorno-Karabakh. SSAR, Salt Lake City, 154 pp. [ISBN: 978-0-916984-84-7.]
  • Arsovski, D. , Olivier, A. , Bonnet, X. , Drilholle, S. , Tomović, L. , Béchet, A. , Golubović, A. and Besnard, A. 2018. Covariates streamline age‐specific early life survival estimates of two chelonian species. J Zool, 306: 223-234. doi:10.1111/jzo.12585 - get paper here
  • Auer, M. 2002. Süsswasserschildkröten in der Westtürkei. Elaphe 10 (3): 34-40
  • AYAZ, D. & ÇIÇEK, K. 2011. Data on the population structure of the European Pond Turtle, Emys orbicularis (LINNAEUS, 1758) in Lake Sülüklü (western Anatolia, Turkey). Herpetozoa 24: - get paper here
  • AYAZ, D. & FRITZ, U. & ATATÜR, M. K. 2007. Morphological comparison of Emys orbicularis (LINNAEUS, 1758) from European Turkey and eastern Bulgaria. Herpetozoa 20 - get paper here
  • AYAZ, D. & TOK, C. V. & ÇIÇEK, K. 2007. Overwintered hatchling of Emys orbicularis (LINNAEUS, 1758) observed in Turkey [Short Note]. Herpetozoa 19 (3/4): 189-192 - get paper here
  • Ayaz, D., U. Fritz, M.K. Atatür, A. Mermer, K. Cicek & M. Afsar 2008. Aspects of Population Structure of the European Pond Turtle (Emys orbicularis) in Lake Yayla, Western Anatolia, Turkey Journal of Herpetology 42 (3): 518-522. - get paper here
  • Ayaz, Dinçer; Kerim Çiçek, Cemal Varol Tok, Hasan Serdar Mutlu, Oğuzkan Cumhuriyet, Yusuf Bayrakcı, Meltem Kumaş. 2013. Conservation activities for European pond turtles (Emys orbicularis) in Turkey. Herpetology Notes 6: 143-144
  • Ayres Fernández, César and Adolfo Cordero Rivera 2001. Sexual dimorphism and morphological differentiation in European Pond Turtle (Emys orbicularis) populations from northwestern Spain. Chelonian Conserv. Biol. 4 (1):100-106
  • Ayres, C. & Iglesias, R. 2017. Emys orbicularis (European Pond Turtle) Ectoparasites. Herpetological Review 48 (4): 834-835 - get paper here
  • Ayres, César; Cordero, Adolfo 2007. Site tenacity in European pond turtle (Emys orbicularis) hatchlings in Northwestern Spain. Amphibia-Reptilia 28 (1): 144-147 - get paper here
  • Ayres-Fernandez, Cesar; Cordero-Rivera, Adolfo 2004. Asymmetries and accessory scutes in Emys orbicularis from Northwest Spain. Biologia, Bratislava 59 (Suppl. 14):85-88.
  • Bakiev, A., Kirillov, A., Kirillova, N., Ruchin, A., Klenina, A., Gorelov, R., & Kostina, N. 2020. Reptile occurrences data in the Volga River basin (Russia). Biodiversity Data Journal, 8, e58033 - get paper here
  • Barroso, David Donaire; Raúl del Canto-Gonzalez, Serge Bogaerts, Frank Pasmans & Hasan el Mouden 2011. Nuevas localidades para Emys orbicularis occidentalis Fritz, 1993 (Testudines: Emydidae) en el Rif (Noroeste de Marruecos). Bull. Soc. Catalana d’Herp. 19 - get paper here
  • Bassu, L.; V. Nulchis, M. G. Satta, C. Fresi & C. Corti 2008. Atlas of amphibians and reptiles of Sardinia - state of the art and general considerations. Herpetologia Sardiniae (Claudia Corti, ed.), Edizioni Belvedere, 504 pp.
  • Bayrakci, Yusef and Dincer Ayaz. 2014. Dynamics of a Central Anatolian population of Emys orbicularis (Linnaeus, 1758) (Testudines: Emydidae). Herpetozoa 27 (1/2): 29-37 - get paper here
  • Bayrakcı, Yusuf; Dinçer Ayaz, Kerim Çiçek 2015. Data on the Population of Syntopic Turtles Emys orbicularis (L., 1758) and Mauremys rivulata (Valenciennes, 1883) from Great Menderes Delta (Western Anatolia, Turkey). Russ. J. Herpetol. 22 (2): 79-83
  • Beau, F. & Brischoux, F. 2017. Emys orbicularis (European Pond Turtle) Neonate diet. Herpetological Review 48 (3): 621-622. - get paper here
  • Bell, T. 1836. A monograph of the Testudinata. London, 1836-1842 (sometimes given as 1832-1836, not date given, unnumbered pages) - get paper here
  • Berroneau, M. et al. 2010. Guide des Amphibiens et Reptiles d’Aquitaine. Association Cistude Nature, 180 pp.
  • Beshkov, V & Nanev, K 2006. The Amphibians and Reptiles in Bulgaria. Pensoft Series Faunistica 46, 120 pp.
  • Bibron & Bory de Saint-Vincent 1833. Vertébrés a sang froid. Reptiles et poissons. Reptiles. In: Geoffroy & Geoffroy, Expédition Scientifique de Morée, Tome III. 1re partie, Zoologie: pp. 57- - get paper here
  • Bidmon, Hans-Jürgen and Ivo E. Ivanchev. 2014. Der südöstliche Zipfel Europas und seine Sumpfschildkröten: Eine Tour auf der Veleka und entlang der Butamjatamündung. Schildkröten im Fokus 11 (2): 21-33
  • Bird, C. G. 1936. The distribution of reptiles and amphibians in Asiatic Turkey with notes on a collection from the Vilayets of Adana, Gaziantep and Malatya. Ann. Mag. Nat. Hist. (10) 18: 257-281. - get paper here
  • Bona, Martin; Adriana Burešová, Stanislav Danko, Peter Havaš, Milan Novotný. 2013. Conservation activities for the European pond turtle (Emys orbicularis) in Slovakia. Herpetology Notes 6: 145-146
  • Bona, Martin; Novotný, Milan; Danko, Stanislav; Burešová, Adriana 2012. Nest site fidelity in the Slovakian population of the European pond turtle Emys orbicularis. Amphibia-Reptilia 33 (2): 207-213 - get paper here
  • Bour, R. 2008. Global diversity of turtles (Chelonii; Reptilia) in freshwater. Hydrobiologia 595:593–598 - get paper here
  • Braitmayer, Nicole;Fritz, Uwe;Mayol, Joan;Pieh, Alexander 1998. Die Europäische Sumpfschildkröte (Emys orbicularis) Menorcas. Elaphe 6 (4): 57-60 - get paper here
  • Brandes, Florian & Martina Anne-Claire Meeske 2015. Ein Wiederansiedlungsprojekt für die Europäische Sumpfschildkröte und die Problematik ausgesetzter Wasserschildkröten aus Privathaltung. Terraria-Elaphe 2015 (3): 32-36 - get paper here
  • Broggi, Mario F. 1978. Herpetologische Beobachtungen auf der Insel Lesbos (Griechenland). Salamandra 14 (4): 161-171 - get paper here
  • Broggi, Mario F. 2018. The “wetland island” of Lemnos (Greece): herpetofauna and nature conservation (Testudines: Amydidae, Geoemydidae; Anura: Pelobatidae). Herpetozoa 30 (3/4): 169 - 178 - get paper here
  • Broggi, Mario F. &; GRILLITSCH, H. 2012. The European Pond Terrapin Emys orbicularis hellenica (VALENCIENNES, 1832), in the Aegean: Distribution and threats. Herpetozoa 25 (1/2): 47-58 - get paper here
  • Cabela, A., & Tiedemann, F. 1985. Atlas der Amphibien und Reptilien Osterreichs. Naturhist. Mus. in Wien, Neue Denkschriften Band 4 80 pp.
  • Calori, Luigi 1853. Sulla matrice degli scudetti cornei della cassa toracico-addominale dei Cheloni. Mem. Acad. Sci. Ist. Bologna 4: 143-166 + 2 plates - get paper here
  • CAPULA, M. & LUISELLI, L. & RUGIERO, L. & FILIPPI, E. 1994. A field experiment on the selection of basking sites by Emys orbicularis (LINNAEUS, 1758). Herpetozoa 7 - get paper here
  • CIOFI, CLAUDIO; ATHANASIA C. TZIKA, CHIARA NATALI,GUIDO CHELAZZI, THEODORUS NAZIRIDIS and MICHEL C. MILINKOVITCH 2009. Characterization of microsatellite loci in the European pond turtle Emys orbicularis. Molecular Ecology Resources 9: 189–191 - get paper here
  • Cogălniceanu, Dan; Laurentiu Rozylowicz, Paul Székely, Ciprian Samoilă, Florina Stănescu, Marian Tudor, Diana Székely, Ruben Iosif 2013. Diversity and distribution of reptiles in Romania. ZooKeys 341 (2013): 49-76 - get paper here
  • Covaciu-Marcov S.-D. et al. 2007. Contributions to the study of the geographical distribution of the herpetofauna in the North-East area of Arad County in Romania. Herpetologica Romanica 1: 62-69
  • Covaciu-Marcov S.-D.; Ghira, I.; Cicort-Lucaciu A.-St., Sas I.; Strugariu, A. & Bogdan H. V. 2006. Contributions to knowledge regarding the geographical distribution of the herpetofauna of Dobrudja, Romania. North-Western Journal of Zoology 2 (2): 88-125 - get paper here
  • ÇAKMAK, Mücahit; Bahadır AKMAN and Mehmet Zülfü YILDIZ 2018. HERPETOFAUNA OF BARTIN PROVINCE (NORTHWEST BLACKSEA REGION, TURKEY). South Western Journal of Horticulture, Biology and Environment P-Issn: 2067- 9874, E-Issn: 2068-7958<br />Vol.8, No.2, 2017 pp.89-102 - get paper here
  • Delaugerre, M., & Cheylan, M. 1992. Atlas de répartition des batraciens et reptiles de Corse. Ajaccio Parc naturel régional de Corse: Ecole pratique des hautes études, 128 pp. - get paper here
  • Díaz-Paniagua,Carmen, Ana Cristina Andreu, Adolfo Marco, Marta Nuez, Judit Hidalgo-Vila, Natividad Perez-Santigosa 2014. Data on nesting, incubation, and hatchling emergence in the two native aquatic turtle species (Emys orbicularis and Mauremys leprosa) from Doñana National Park. Basic and Applied Herpetology 28: 145-151 - get paper here
  • Dietrich, N. 2008. ...und wieder mal am Goldstrand. Elaphe 16 (2): 50-53
  • Drechsler, Robby M.; Lucía M. Fernández, Matthieu Lassalle, and Juan S. Monrós 2018. Movement Patterns and Home Range Sizes of Translocated European Pond Turtles (Emys orbicularis, Linnaeus, 1758) in Eastern Spain. Herp. Cons. Biol. 13 (1): 1–9 - get paper here
  • Dubois, A. & Bour, R. 2010. The nomenclatural status of the nomina of amphibians and reptiles created by Garsault (1764), with a parsimonious solution to an old nomenclatural problem regarding the genus Bufo (Amphibia, Anura), comments on the taxonomy of this genus, and comment Zootaxa 2447: 1–52 - get paper here
  • Duméril, A. M.C., G. BIBRON & A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 9. Paris, XX + 440 S. - get paper here
  • Duméril, A.M.C., and G. Bibron. 1835. Erpétologie Générale ou Histoire Naturelle Complète des Reptiles, Vol. 2. Librairie Encyclopédique de Roret, Paris, iv + 680 p. - get paper here
  • Dunaev E.A., Orlova V.F. 2017. Amphibians and reptiles of Russia. Atlas and determination. 2nd. ed. Moscow: Phyton XXI, 328 p
  • Dux, Marta; Daniel Doktór, Adam Hryniewicz& Beata Prusak 2017. Evaluation of 11 Microsatellite Loci for Reconstructing of Kinship Groups in the European Pond Turtle, Emys orbicularis (Linnaeus, 1758). Acta zool. bulg., Suppl. 10, 2017: 15-22 - get paper here
  • Duysebaeva, T. N.; I. V. Doronin, D. V. Malakhov, O. V. Kukushkin, A. G. Bakiev <br /> 2019. GIS analysis of the distribution and habitation conditions of Emys orbicularis оrbicularis (Testudines, Emydidae): methodical aspects. PROCEEDINGS OF THE VII CONGRESS OF THE NIKOLSKY HERPETOLOGICAL SOCIETY, in "University proceedings. Volga region. Natural sciences” 1: 28-40 - get paper here
  • Egretzberger,G. 1995. Taxonomie aktuell [list of subspecies, their distribution + map]. Informationsblatt der Fachgruppe Schildkröten (ÖGH) 1 (1): 8-11
  • Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Erdélyi G, Szabó B, Kiss I 2019. Basking activity pattern of the European pond turtle, Emys orbicularis (Linnaeus, 1758) in Babat valley (Gödöllő, Hungary). Herpetozoa 32: 221-227 - get paper here
  • Erdélyi, Gergő; Borbála Szabó, István Kiss 2021. Basking Site Selection and Usage Strategies of the European Pond Turtle, Emys orbicularis (Linnaeus, 1758) in Babat valley (Gödöllő, Hungary). Russian Journal of Herpetology 28 (4): 175-184 - get paper here
  • Ernst,C.H. and Barbour,R.W. 1989. Turtles of the World. Smithsonian Institution Press, Washington D.C. - London
  • Escoriza, Daniel; Marc Franch, Santiago Ramos, Pau Sunyer-Sala, and Dani Boix 2020. Demographics and Survivorship in the European Pond Turtle (Emys orbicularis): a 31–Year Study. Herpetological Conservation and Biology, 15 (1): - get paper here
  • Fahd, S.; el Marnisi, B.; Mediani, M. & Fritz, U. 2009. Zur Verbreitung und zurm Bedrohungsstatus der Europäischen Sumpfschildkröte (Emys orbicularis) in Marokko. Elaphe 17 (3): 30-33 - get paper here
  • Farkas, Balázs; Bálint Halpern, Péter Agócs, Róbert Dankovics, Angéla Földi, Erika Gulácsi, György Györffy, Zsófia Kalmár, István Kiss, Tibor Kovács, Zsófia Eszter Lovász, László Molnár, Tamás Gergely Molnár, Tamás Péchy, Tibor Somlai, László Torvaji 2013. Conservation activities for European pond turtles (Emys orbicularis) in Hungary. Herpetology Notes 6: 107-110
  • Fediras, Samira; Rachid Rouag, Nadia Ziane, Anthony Olivier, Arnaud Béchet, Slim Benyacoub 2018. Population Structure and Morphometrics of the European Pond Turtle (Emys orbicularis (Linnaeus, 1758)) at Lake Tonga, Algeria. Russ. J. Herpetol. 25 (2): 88-96 - get paper here
  • Fernández-Ortín, Daniel; Gregorio Sánchez-Montes, Íñigo Martínez-Solano 2019. A catalogue of reptiles of Monfragüe National Park (Spain), with molecular characterization of populations of Blanus WAGLER, 1830 in this protected area. Basic and Applied Herpetology - get paper here
  • Ficetola, Gentile Francesco; De Bernardi, Fiorenza 2006. Is the European "pond" turtle Emys orbicularis strictly aquatic and carnivorous? Amphibia-Reptilia 27 (3): 445-447 - get paper here
  • Ficetola, Gentile Francesco; Sebastiano Salvidio, Stefania D’Angelo, Anna Bonardi, Luciana Bottoni, Laura Canalis, Stefano Crosetto, Silvia Di Martino, Vincenzo Ferri, Paolo Filetto, Paolo Genta, Riccardo Jesu, Simone Masin, Stefano Mazzotti, Dario O 2013. Conservation activities for European and Sicilian pond turtles (Emys orbicularis and Emys trinacris, respectively) in Italy. Herpetology Notes 6: 127-133
  • Forcart, Lothar 1950. Amphibien und Reptilien von Iran. Verh. naturf. Ges. Basel 61: 141-156
  • Frank, Thomas; Enerit Saçdanaku, Michael Duda & Ferdinand Bego 2018. Amphibian and reptile fauna of the Vjosa River, Albania. Acta ZooBot Austria 155: 323–336 - get paper here
  • Freund, Michael 2013. Die Sumpfschildkröte (Emys orbicularis) im Biologieunterricht. Terraria-Elaphe 2013 (4): 29-38 - get paper here
  • Fritz, U. 1989. Zur innerartlichen Variabilität von Emys orbicularis (LINNAEUS 1758). 1. Eine neue Unterart der Europäischen Sumpfschildkröte aus Kleinasien, Emys orbicularis luteofusca subsp. nov. Salamandra 25 (3-4): 143-168 - get paper here
  • FRITZ, U. 1993. Weitere Mitteilung zur innerartlichen Variabilität, Chorologie und Zoogeographie von Emys orbicularis (LINNAEUS, 1758) in Kleinasien. Herpetozoa 6 - get paper here
  • Fritz, U. 1994. Gibt es in Nordafrika zwei verschiedene Formen der Europäischen Sumpfschildkröte (Emys orbicularis)? Salamandra 30 (1): 76-80 - get paper here
  • Fritz, U. 1994. Zur innerartlichen Variabilität von Emys orbicularis (LINNAEUS, 1758). 4. Variabilität und Zoogeographie im pontokaspischen Gebiet mit Beschreibung von drei neuen Unterarten (Reptilia; Testudines: Emydidae). Zool. Abh. Mus. Tierk. Dresden, 48 (4): 53-93. - get paper here
  • Fritz, U. 1995. Kritische Übersicht der Fossilgeschichte der Sumpfschildkröten-Gattung Emys A. Duméril, 1806. Zool. Abh. (Dresden) 48 (3): 243-264. - get paper here
  • Fritz, U. 1995. Zur innerartlichen Variabilität von Emys orbicularis (LINNAEUS, 1758). 5a. Taxonomie in Mittel-Westeuropa , auf Korsika, Sardinien, der Apenninen-Halbinsel und Sizilien und Unterartengruppen von Emys orbicularis (Reptilia: Testudines: Emydidae). Zool. Abh. Mus. Tierk., Dresden, 48 (3): 185-242.
  • Fritz, U. 2003. Die Europäische Sumpfschildkröte (Emys orbicularis). Laurenti Verlag (Bielefeld), Zeitschrift für Feldherpetologie, Suppl. 1, 224 pp. - get paper here
  • Fritz, U., C. KELLER & M. BUDDE 1996. Eine neue Unterart der Europäischen Sumpfschildkröte aus Südwestspanien, Emys orbicularis hispanica subsp. nov. Salamandra 32 (3): 129-152 - get paper here
  • Fritz, Uwe 1995. Die Sumpfschildkröte in Oberschwaben oder: Die Suche nach einem Phantom? Elaphe 3 (4): 57-61 - get paper here
  • Fritz, Uwe, Daniela Guicking, Hagi Gholi Kami, Marine Arakelyan, Markus Auer, Dínçer Ayaz, Ceesar Ayres Fernández, Andrey G. Bakiev, Antonia Celani, Georg Dzukic, Soumia Fahd, Peter Havas, Ulrich Joger, Viner Khabibullin, Lyudmila Mazanaeva, Pavel Si 2007. Mitochondrial phylogeography of European pond turtles Emys orbicularis, Emys trinacris - an update. Amphibia-Reptilia 28 (3): 418-426 - get paper here
  • Fritz, Uwe. 1998. Introduction to zoogeography and subspecific differentiation in Emys orbicularis (Linnaeus, 1758). Mertensiella. 10: 1-27 - get paper here
  • Fritz, Uwe; Cadi, Antoine; Cheylan, Marc; Coïc, Christophe; Détaint, Mathieu; Olivier, Anthony; Rosecchi, Elisabeth; Guicking, Daniela; Lenk, Peter; Joger, Ulrich; Wink, Michael 2005. Distribution of mtDNA haplotypes (cyt b) of Emys orbicularis in France and implications for postglacial recolonization. Amphibia-Reptilia 26 (2): 231-238 - get paper here
  • Fritz, Uwe; Dinçer Ayaz, Anna K. Hundsdörfer, Tatiana Kotenko, Daniela Guicking, Michael Wink, C. Varol Tok, Kerim Çiçek, Jutta Buschbom 2009. Mitochondrial diversity of European pond turtles (Emys orbicularis) in Anatolia and the Ponto-Caspian Region: Multiple old refuges, hotspot of extant diversification and critically endangered endemics. Organisms Diversity & Evolution 9 (2): 100-114 - get paper here
  • Fritz, Uwe; Ibrahim Baran; Abidin Budak and Eike Amthauer 1998. Some notes on the morphology of Emys orbicularis in Anatolia, especially on E. o. luteofusca and E. o. colchica, with the description of a new subspecies from southeastern Turkey. Mertensiella 10: 103-121. - get paper here
  • Fritz, Uwe; Kuzmin, Sergius L.; Kolobajewa, Olga W.; Orlowa, Walentina F. 1995. Zur Variabilität der Sumpfschildkröte (Emys orbicularis) im Gebiet zwischen der Manytsch-Niederung und Don (Russland). Salamandra 31 (4): 231-236 - get paper here
  • Fritz, Uwe;Obst, Fritz Jürgen 1995. Morphologische Variabilität in der Intergradationzonen von Emys orbicularis orbicularis und E. o. hellenica. Salamandra 31 (3): 157-180 - get paper here
  • Frynta,D. et al. 1997. Results of the Czech biological expedition to Iran. Part 1. Notes on the distribution of amphibians and reptiles. Acta Soc. Zool. Bohem. 61: 3-17
  • GARIBOLDI, A. & ZUFFI, A. L. 1994. Notes on the population reinforcement project for Emys orbicularis (LINNAEUS, 1758) in a natural park of northwestern Italy. Herpetozoa 7 - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Gent, Tony G 2013. Historical and current situation of the European pond turtle (Emys orbicularis) in the United Kingdom. Herpetology Notes 6: 117-118
  • Gholamifard A, Esmaeili HR, Rastegar-Pouyani N. 2015. A noteworthy record of translocation for Emys orbicularis persica, Eichwald 1831, in southern Iran. Amphibian & Reptile Conservation 9 (1): 49-53 - get paper here
  • Girondot M, Zaborski P, Servan J, Pieau C. 1994. Genetic contribution to sex determination in turtles with environmental sex determination. Genet Res 63:117–127. - get paper here
  • Girondot, Marc; Servan, Jean; Pieau, Claude 2011. Rapid search for genetic differentiation of populations: application to the European pond turtle. Amphibia-Reptilia 32 (3): 297-304 - get paper here
  • Gläss, H. & Meusel, W. 1996. Die Süsswasserschildkröten Europas. Neue Brehm-Bücherei Bd. 418, Westarp/Spektrum, 80 pp.
  • Gramentz, D. 1993. Neuer Fundort von Emys orbicularis (Linnaeus, 1758) im Südwesten der Türkei. Herpetofauna 15 (82): 06-08 - get paper here
  • Gray,J.E. 1856. On some new species freshwater tortoises from North America, Ceylon and Australia. Ann. Mag. Nat. Hist. (2) 18: 263-268 - get paper here
  • Haxhiu, Idriz 1995. Results of studies on the chelonians of Albania. Chelonian Conserv. Biol. 1 (4): 324-326
  • Henle, K. 1985. Ökologische, zoogeographische und systematische Bemerkungen zur Herpetofauna Jugoslawiens. Salamandra 21 (4): 229-251 - get paper here
  • Hennig, A.S. 2004. Haltung von Wasserschildkröten. Natur und Tier Verlag (Münster), 125 pp. - get paper here
  • Hennig, Andreas S. 2013. Die Neuwelt-Sumpfschildkröten der Familie Emydidae. Terraria-Elaphe 2013 (4): 14-20 - get paper here
  • Hiller, A. 1990. Eine Europäische Sumpfschildkröte Emys orbicularis (LINNAEUS, 1758) mit angeborener letaler Mißbildung. Salamandra 26 (2-3): 215-217 - get paper here
  • Hofstra, Jelle 2015. Sumpfschildkröten auf der griechischen Insel Lesbos. Schildkröten im Fokus. 12 (3): 27-35
  • Horváth, E., Martvoňová, M., Danko, S., Havaš, P., Kaňuch, P., & Uhrin, M. 2021. Distribution range and population viability of Emys orbicularis in Slovakia: a review with conservation implications. Nature Conservation, 44, 141
  • Horváth, Enikő; Peter Kaňuch, Marcel Uhrin 2021. Predation on nests of the European pond turtle (Emys orbicularis): remarks from failed field experiments. Herpetology Notes 14: 1067-1072
  • HOSSEINIAN-YOUSEFKHANI, SEYYED SAEED; AMAAL GH. YASSER , MURTADA D. NASER, & ESKANDAR RSATEGAR-POUYANI 2021. Increased haplotype diversity of Emys orbicularis (Linnaeus, 1758) (Reptilia: Emydidae) in northern Iran. Ecologica Montenegrina 45: 24-32 - get paper here
  • Hughes, B. 1986. Longevity Records of African Captive Amphibians and Reptiles: Part 1: Introduction and Species List 1 - Amphibians and Chelonians. J. Herp. Assoc. Africa (32): 1-5 - get paper here
  • Isailović, Jelka Crnobrnja; Gabor Mesaroš. 2013. Conservation activities for the European pond turtle (Emys orbicularis) in Serbia. Herpetology Notes 6: 119-121
  • Jablonski, Daniel 2011. Reptiles and amphibians of Albania with new records and notes on occurrence and distribution. Acta Soc. Zool. Bohem. 75: 223–238 - get paper here
  • Jablonski, Daniel; Jakub Mrocek, Daniel Gruľa, Jana Christophoryová 2017. Attempting courtship between Emys orbicularis and Trachemys scripta (Testudines: Emydidae). Herpetology Notes 10: 123-126 - get paper here
  • Jamniczky, H. A. and A. P. Russell 2007. Reappraisal of Patterns of Nonmarine Cryptodiran Turtle Carotid Circulation: Evidence From Osteological Correlates and Soft Tissues. Journal of Morphology 268:571–587 - get paper here
  • Jendretzke, N. 2011. La Brenne, ein Paradies für Europäische Sumpfschildkröten in Frankreich. Elaphe 19 (2): 20-24
  • Jesu R., Piombo R., Salvidio S., Lamagni L., Ortale S., Genta P. 2004. Un nuovo taxon di testuggine palustre endemico della Liguria Occidentale. Ann. Mus. Civ. St. Nat. “G. Doria” Genova. - get paper here
  • Joger, U.; Fritz, U.; Guicking, D.; Kalyabina-Hauf, S.; Nagy, Z.T. & Wink, M. 2007. Phylogeography of western Palaearctic reptiles – Spatial and temporal speciation patterns. Zoologischer Anzeiger 246: 293–313 - get paper here
  • Joyce, Walter G.; James F. Parham , Tyler R. Lyson , Rachel C. M. Warnock and Philip C. J. Donoghue 2013. A Divergence Dating Analysis of Turtles Using Fossil Calibrations: An Example of Best Practices. Journal of Paleontology, 87(4):612-634 - get paper here
  • Kabisch, K. 1966. Herpetologische Exkursionen in die Umgebung von Sofia. Aquarien-Terrarien, 4:131-135
  • Kabisch, K. & Belter, H. 1968. Das Verzehren von Amphibien und Reptilien durch Vögel. Zoologische Abhandlungen aus dem Staatlichen Museum für Tierkunde in Dresden 29: 191-227
  • Kabisch, K. & Engelhard, H. 1964. Beobachtungen an Reptilien in der Umgebung von Varna-Goldsand. Aquarien Terrarien 11 (6): 194-197
  • Kabisch, K. & Rogner, M. 1988. Eineiige Zwillinge der Europäischen Sumpfschildkröte, Emys orbicularis (L.). Aquarien Terrarien 35: 416-417
  • Kasapidis, Panagiotis;Provatidou, Sofia;Maragou, Panagiota;Valakos, Efstratios D. 1996. Neue Daten über die Herpetofauna von Lesbos (ägäische Inseln, Griechenland) und einige biogeographische Bemerkungen über die Inseln des nordöstlichen ägäischen Archipels. Salamandra 32 (3): 171-180 - get paper here
  • Kleewein, Andreas and Günther Wöss. 2013. Status of the European Pond Turtle, Emys orbicularis (Reptilia: Testudines: Emydidae) in Vorarlberg, Austria. Acta Herpetologica 8 (1): 65-67
  • Kleewein, Andreas. 2016. First genetic investigations into the occurrence of Emys orbicularis (LINNAEUS, 1758) in Carinthia, Austria. Herpetozoa 29 (1/2): 90-92 - get paper here
  • Kuzmin, S.L. 2002. The turtles of Russia and other Ex-soviet Republics. Edition Chimaira, Frankfurt, 159 pp.
  • Kwet, A. 2010. Reptilien und Amphibien Europas. Kosmos-Verlag, Stuttgart, 252 pp.
  • Kwet, Axel & Benny Trapp 2014. Liste der Reptilien Europas. Draco 15 (60): 72-79 - get paper here
  • Kwet, Axel; Uwe Fritz, Andreas Nöllert, Richard Podloucky & Norbert Schneeweiss 2015. Von der Fastenspeise zum Reptil des Jahres 2015 – die Europäische Sumpfschildkröte. Terraria-Elaphe 2015 (3): 14-23 - get paper here
  • Laufer, H.; Fritz, C. & Sowig, P. (eds.) 2007. Die Amphibien und Reptilien Baden-Württembergs. Ulmer Verlag, Stuttgart, 807 pp.
  • Lebboroni, Marco; Cecchini, Agnese 2005. Basking Counts as Abundance Indices in Pond Populations of Emys Orbicularis. The Herpetological Journal 15: 121-124 - get paper here
  • Lehmann, K. 1980. Zum Vorkommen der Europäischen Sumpfschildkröte, Emys orbicularis, auf Menorca/Balearen (Spanien) (Reptilia: Testudines: Emydidae). Salamandra 16 (2): 132-134 - get paper here
  • Leidy, J. 1888. Entozoa in the Terapin. Proc. Acad. Nat. Sci. Philadelphia 1888: 127-128 - get paper here
  • Lenk, P., Fritz, U., Joger, U., and Wink, M. 1999. Mitochondrial phylogeography of the European pond turtle, Emys orbicularis (Linnaeus 1758). Molecular Ecology 8: 1911–1922. - get paper here
  • Leptien, R. 2013. Die Europäische Sumpfschildkröte. Reptilia (Münster) 18 (100): 97-100 - get paper here
  • Lever, C. 2003. Naturalized reptiles and amphibians of the world. Oxford University Press, 338 pp.
  • Link, H.F. 1807. Beschreibung der Naturalien-Sammlung der Universität zu Rostock, zweite Abtheilung. Adlers Erben, Rostock, pp. 51-100 - get paper here
  • Linnaeus, C. 1758. Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Holmiæ. 10th Edition: 824 pp. - get paper here
  • Lourenço, João M.; Julien Claude, Nicolas Galtier, Ylenia Chiari 2011. Dating cryptodiran nodes: Origin and diversification of the turtle superfamily Testudinoidea. Molecular Phylogenetics and Evolution 62 (1): - get paper here
  • Mähn, M. 2003. Die Europäische Sumpfschildkröte - Emys orbicularis (LINNAEUS 1759) - die ideale Schildkröte für den Gartenteich. Draco 4 (13): 19-24 - get paper here
  • Malkmus, R. 1982. Beitrag zur Verbreitung der Amphibien und Reptilien in Portugal. Salamandra 18 (3-4): 218-299 - get paper here
  • Malkmus, R. 1995. Die Amphibien und Reptilien Portugals, Madeiras und der Azoren. Die Neue Brehm-Bücherei, Bd. 621. Westarp Wissenschaften (Magdeburg)/Spektrum Akademischer Verlag (Heidelberg)
  • Martens, Hans-Joachim. 2016. Observations of the European Pond Turtle. Radiata (English edition) 25 (1): 25-27
  • Martens, Hans-Joachim. 2016. Beobachtungen zur Europäischen Sumpfschildkröte. Radiata (English edition) 25 (1): 25-27
  • Martínez del Mármol, Gabriel; D. James Harris, Philippe Geniez, Philip de Pous, and Daniele Salvi 2019. Amphibians and Reptiles of Morocco. Edition Chimaira, Frankfurt am Main, Germany, 478 pp - get paper here
  • Mascort, Ramón, Albert Bertolero and Oscar J. Arribas 1999. Morphology, geographic variation and taxonomy of Emys orbicularis L. 1758, in the northeast of the Iberian Peninsula. Revista Española de Herpetología 13: 7-16. - get paper here
  • Meeske, A.-C. M. & E. Snieshkus & F. W. Welter-Schultes 2000. Aufzucht und Wachstumsanalysen der Europäischen Sumpfschildkröte (Emys orbicularis) aus Litauen. Salamandra 36 (2): 89-102 - get paper here
  • Meeske, Anne Clarie Martina, Mihails Pupins and Krzysztof Rybczynski 2006. Erste Ergebnisse zur Verbreitung und zum Status der Europäischen Sumpfschildkröte (Emys orbicularis) am nördlichen Rand ihrer Verbreitung in Litauen und Lettland. Zeitschrift für Feldherpetologie 13 (1):71-99. - get paper here
  • Meiri, Shai; Amos Belmaker, Daniel Berkowic, Kesem Kazes, Erez Maza, Guy Bar-Oz and Roi Dor 2019. A checklist of Israeli land vertebrates. Israel Journal of Ecology and Evolution - get paper here
  • Melita, Vamberger; Poboljšaj Katja, Marijan Govedič, Nika Debeljak Šabec, Anamarija Žagar. 2013. Conservation activities for European pond turtles (Emys orbicularis) in Slovenia. Herpetology Notes 6: 123-126
  • Mertens, Robert. 1950. Einige Beobachtungen bei der Zucht der Europäischen Sumpfschildkröte. Der Zoologische Garten 17: 170-175
  • Methner, K. 1988. Zum Vorkommen von Emys orbicularis (LINNAEUS 1758) auf Menorca (Balearen) II. Salamandra 24 (4): 302-305 - get paper here
  • Mieza-Paez, Eduardo, Diego Martínez-Martínez, Aida Tarragó, Angel Such Sanz, Josep Nogués Vera 2016. Current situation and environmental factors affecting the distribution of Emys orbicularis in Sèquia Major (NE Iberian Peninsula) in syntopy with Mauremys leprosa. Basic and Applied Herpetology 30: 47-59 - get paper here
  • Mignet, Fabien; Jean-Yves Barnagaud, Laurent Barthe, Albert Bertolero, Valérie Bosc, Florine Escot, Romain Fleuriau, Laurent Joubert, André Joyeux, Damien Lerat, Grégoire Massez, Anthony Olivier, Zoey Owen-Jones, Kathleen Perrot, Jérémiah Petit, Fran 2020. Geographic variation in body size among French populations of the European pond turtle. Amphibia-Reptilia 42 (2): 141–151 [2021] - get paper here
  • Mignet, Fabien; Thomas Gendre, Denis Reudet, Frédérique Malgoire, Marc Cheylan, and Aurélien Besnard 2014. Short-Term Evaluation of the Success of a Reintroduction Program of the European Pond Turtle: The Contribution of Space-Use Modeling. Chelonian Conservation and Biology Jul 2014, Vol. 13, No. 1: 72-80. - get paper here
  • Mitrus, Sawomir 2005. Headstarting in European pond turtles (Emys orbicularis): Does it work? Amphibia-Reptilia 26 (3): 333-341 - get paper here
  • Mitrus, Slawomir; Najbar, Bartlomiej; Kotowicz, Adam 2012. Frequency of successful reproduction and time of nest emergence of hatchlings of the European pond turtle in the northern part of its distribution area. The Herpetological Journal 22 (4): 235-239 - get paper here
  • Mitrus, Sławomir 2009. Growth rings in young turtles Emys orbicularis – marking is the only reliable criterion for distinguishing between wild and headstarted animals. The Herpetological Journal 19: 107-109 - get paper here
  • Mitrus, Sławomir 2010. Is the European pond turtle Emys orbicularis strictly aquatic? – Habitats where the turtle lives in central Europe. Acta Herpetologica 5 (1): 31-35 - get paper here
  • Mitrus, Sławomir;Bartłomiej Najbar,Adam Kotowicz,Anna Najbar 2018. The Body Size of Headstarted and Wild Juvenile European Pond Turtles (Emys orbicularis). Russ. J. Herpetol. 25 (2): 161-164 - get paper here
  • Mollov, Ivelin A.; Borislav Y. Naumov, Dobrin D. Dobrev, Georgi S. Popgeorgiev. 2013. Conservation activities for European pond turtles (Emys orbicularis) in Bulgaria. Herpetology Notes 6: 135-138
  • Molnár, Tamás; Lehoczky, István; Molnár, Marcell; Benedek, Ildikó; Magyary, István; Jeney, Zsigmond; Zsolnai, Attila 2011. Genetic diversity of the European pond turtle (Emys orbicularis) in the South-West region of Hungary - first results. Amphibia-Reptilia 32 (4): 519-526 - get paper here
  • Moravec J. 2003. Some notes on the population of dwarfed Emys orbicularis from Pag (Croatia). Journal of the National Museum (Prague), Natural History Series 172 (1-4): 55-60
  • Mutz, T.; Mutz, M. & Obst, F.-J. 1999. Herpetologische Impressionen von Sardinien. Elaphe 7 (3): 76-80
  • Najbar, Bartlomiej & Ewa Szuszkiewicz 2006. The morphometrics and colouration of the European pond turtle Emys orbicularis in Lubuskie province (West Poland). Biologia, Bratislava, 61/5: 585—592 - get paper here
  • NAUMOV, BORISLAV; NIKOLAY TZANKOV, KRASIMIR DONCHEV, BOYAN PETROV, ANDREI STOJANOV, GEORGI POPGEORGIEV, IVELIN MOLLOV, VLADIMIR BESHKOV 2016. The Herpetofauna (Amphibia and Reptilia) of Vrachanska Planina Mountains - Species Composition, Distribution and Conservation. In: Bechev, D. & Georgiev, D. (Eds.), Faunistic diversity of Vrachanski Balkan Nature Park. ZooNotes, Supplement 3, Plovdiv University Press, Plovdiv - get paper here
  • Necas, Petr; Modry, David & Zavadil, Vít 1997. Czech Recent and Fossil Amphibians and Reptiles. Edition Chimaira, Frankfurt, 94 pp.
  • Nekrasova, O.; O. Marushchak, O. Oskyrko 2019. To the study of herpetofauna of "Yelanetskyi steppe" nature reserve and adjoined territоries [in Russian]. Bull. Taras Shevchenko Nat Univ (Biol) 78 (2): 35-40 - get paper here
  • Nöllert, A.; Nöllert, C. & Ritter, A. 1986. Einige Beobachtungen zur Herpetofauna der bulgarischen Schwarzmeerküste und Südwestbulgariens (Teil 2 - Die Reptilien). Herpetofauna 8 (44): 30-34 - get paper here
  • Ottonello, Dario; Fabrizio Oneto, Monica Vignone, Anita Rizzo, Sebastiano Salvidio 2018. Diet of a restocked population of the European pond turtle Emys orbicularis in NW Italy Acta Herpetologica 13 (1): 89-93 - get paper here
  • Paepke, Hans-Joachim 2007. Prof. Dr. h.c. Bruno Dürigen, his life and his achievements in the fields of herpetology, vivarium- and poultry science. Mitteilungen aus dem Museum für Naturkunde in Berlin 83 (S1): 74-82 - get paper here
  • Pawlowski, S. 2005. Vergleichende Ei-Entwicklungsdauer der Europäischen Sumpfschildkröte, Emys orbicularis (LINNAEUS 1758) und der Kaspischen Flussschildkröte, Mauremys rivulata (VALENCIENNES 1833). Elaphe 13 (3): 29-33
  • Pawlowski, S. 2009. Freilandschlupf bei der Europäischen Sumpfschildkröte, Emys orbicularis ssp. LINNAEUS , 1758, aus einem im Freiland überwinterten Gelege. Elaphe 17 (2): 22-30
  • PAWLOWSKI, S. 2015. Ergänzende Bemerkungen zur Überwinterung verschiedener aquatischer Schildkröten unter kontrollierten Bedingungen. Sauria 37 (2): 58 - 63 - get paper here
  • PAWLOWSKI, S. 2016. Notes on the outdoor husbandry and breeding of the French Pond Turtle, Emys orbicularis galloitalica Fritz, 1995. Sauria 38 (2): 47 - 53 - get paper here
  • PAWLOWSKI, S. 2017. Präzision bei der Eiablage einer hellenischen Sumpfschildkröte Emys orbicularis hellenica (Valenciennes in Bibron & Bory de Saint-Vincent, 1832). Sauria 39 (4): 19-23 - get paper here
  • PAWLOWSKI, S. 2019. Bemerkungen zur Reproduktion der Ostmediterranen Sumpfschildkröte Emys orbicularis hellenica (Valenciennes, 1832) unter kontrollierten Freilandbedingungen in Mitteleuropa. Sauria 41 (3): 9-18 - get paper here
  • PAWLOWSKI, S. 2020. Bemerkungen zur Reproduktion der Europäischen Sumpfschildkröte Emys orbicularis orbicularis Linnaeus, 1758 unter kontrollierten Freilandbedingungen in Mitteleuropa. Sauria 42 (2): 37-44
  • Pawlowski, S. & Krämer, C. 2009. Bemerkungen zur Herpetologie des nordwestlichen Kroatiens. Sauria 31 (4): 19-30 - get paper here
  • Pedall, I.; Schäfer, H.; Fritz, U. & Wink, M. 2008. Isolation of microsatellite markers in the Emys orbicularis complex and development of multiplex PCR amplification. Conservation Genetics DOI 10.1007/s10592-008-9631-0 - get paper here
  • Pereira, P, Teixeira, J, Velo‐Antón, G. 2018. Allele surfing shaped the genetic structure of the European pond turtle via colonization and population expansion across the Iberian Peninsula from Africa. J Biogeogr. 45: 2202– 2215 - get paper here
  • Perianes, Mario J., Juan M. D. B. Gómez, Maria D. Fernandez-Alzas, Jose L. Perez-Bote, Antonio M. d. Viejo and Manuel Blasco. 2006. Aplicaciones del analisis de imagen como herramienta taxonomica en la discriminacion de poblaciones de Emys orbicularis (L. 1758). Boletin de la Asociacion Herpetologica Española 17(2):109-113 - get paper here
  • Petrov B. P., N. Tzankov, H. Strijbosch, G. Popgeorgiev, V. Beshkov 2006. The herpetofauna (Amphibia and Reptilia) of the Western Rhodopes mountain (Bulgaria and Greece). In: Beron P. (ed.). Biodiversity of Bulgaria. 3. Biodiversity of Western Rhodopes (Bulgaria and Greece) I. Pensoft & Nat. Mus. Natur. Hist., Sofia, pp. 863-912
  • Pfau, J. 1988. Beitrag zur Verbreitung der Herpetofauna in der Niederalgarve (Portugal). Salamandra 24 (4): 258-275 - get paper here
  • Pieau C. 1998. Temperature-dependent sex determination in Emys orbicularis: laboratory and field studies. Mertensiella 10: 199-207
  • Pottier G., Paumier J.-M., Tessier M., Barascud Y., Talhoët S., Liozon R., D’Andurain P., Vacher J.-P., Barthe L., Heaulmé V., Esslinger M., Arthur C.-P., Calvet A., Maurel C. & Redon H. 2008. Atlas de répartition des reptiles et amphibiens de Midi-Pyrénées. Les atlas naturalistes de Midi-Pyrénées. Nature Midi-Pyrénées, Toulouse, 126 pp.
  • Prusak, Beata, Slawomir Mitrus, Bartlomiej Najbar, Alicja Pacholewska, Adrian Deas, Katarzyna Skonieczna, Grzegorz Gorecki, Grzegorz Grzybowski, Adam Hryniewicz, Anton Wroblewski and Tomasz Grzybowski. 2013. Population differentiation of the European Pond Turtle (Emys orbicularis) in Poland inferred by the analysis of mitochondrial and microsatellite DNA: implications for conservation. Amphibia-Reptilia 34 (4): 451-461 - get paper here
  • Raemy, Matthieu; Jean-Claude Monney, Sylvain Ursenbacher. 2013. Conservation activities for European pond turtles (Emys orbicularis) in Switzerland. Herpetology Notes 6: 111-112
  • Rakauskas, Vytautas; Rūta Masiulytė, Alma Pikūnienė 2016. Predator-prey interactions between a recent invader, the Chinese sleeper (Perccottus glenii) and the European pond turtle (Emys orbicularis): a case study from Lithuania. Acta Herpetologica 11 (2): 101-109 - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Rathke,H. 1846. On the development of the Chelonians. Ann. Mag. Nat. Hist. (1) 18: 316-323 - get paper here
  • Rogner, M. 2013. Die Schildkröten von Kalogria (Peloponnes). Reptilia (Münster) 18 (99): 76-78 - get paper here
  • Romanova E. B., Stolyarova I. A., Bakiev A. G., Gorelov R. A. 2021. Comparative leukocyte blood profile of Emys orbicularis (Reptilia: Emydidae) from two populations. Current Studies 21 (1-2): 30 - get paper here
  • Rovero, F., M. Lebboroni & G. Chelazzi 1999. Aggressive Interactions and Mating in Wild Populations of the European Pond Turtle Emys orbicularis Journal of Herpetology 33 (2): 258-263. - get paper here
  • Rutschke, J.; Koepe, D. & Deichsel, G. 2005. Beobachtungen zu anthropogenen Einflüssen auf die Reptilienfauna des Peloponnes (Griechenland). Herpetofauna 25 (143): 17-28 [2003/2004] - get paper here
  • Šalamon, Dragica; Biljana Janev Hutinec, Mila Lončar. 2013. Conservation activities for European pond turtles (Emys orbicularis) in Croatia. Herpetology Notes 6: 149-152
  • Salvi, Daniele; Pierluigi Bombi 2010. Reptiles of Sardinia: updating the knowledge on their distribution. Acta Herpetologica 5 (2): 161-178 - get paper here
  • Schleich, H.H., Kästle,W., Kabisch, K. 1996. Amphibians and Reptiles of North Africa. Koeltz, Koenigstein, 627 pp.
  • Schlüter, U. 2005. Die Herpetofauna der bulgarischen Schwarzmeerküste - Teil 1: Naturraum und Schildkröten. Elaphe 13 (3): 48-56
  • Schmidt B, Kranželić D, Majetić Đ, Lauš B, Štih A, Koren T 2020. Distribution and conservation status of the herpetofauna of Dugi Otok Island, Croatia. Herpetozoa 33: 191-205 - get paper here
  • Schneeweiß, Norbert 2006. On the status of Emys orbicularis on Zakynthos Island. . Radiata 15(2): 30-31.
  • Schneeweiß, Norbert and Hartmut Breu. 2013. Conservation activities for the European pond turtle (Emys orbicularis) in Germany. Herpetology Notes 6: 113-115
  • Schneider,B. 1986. Zur Herpetofauna der Insel Limnos (Hellespontische Inseln, Nordägäis, Griechenland). Salamandra 22 (4): 276-280 - get paper here
  • Schreitmüller, W. 1923. Beiträge zum Verbreitungsgebiet der Sumpfschildkröte (= Emys orbicularis L.) in der Umgebung von Dresden und anderen Orten Sachsens. Archiv für Naturgeschichte 89A (1): 152-160
  • Schulze, Anja and Uwe Fritz 2003. Morphological variation in Tyrrhenian Emys orbicularis revisited. Amphibia-Reptilia 24 (2): 230-234 - get paper here
  • Sehnal, P. & Schuster, A. 1999. Herpetologische Beobachtungen auf der Kvarnerinsel Cres, Kroatien. Ergebnisse von fünf Exkursionen. Herpetozoa 12 (3-4): 163-178 - get paper here
  • Seidel, Michael E. & Carl H. Ernst 2017. A Systematic Review of the Turtle Family Emydidae. Vertebrate Zoology 67 (1): 1–122 - get paper here
  • Shammakov, S. 1981. Presmykaiushchiesia ravninnogo Turkmenistana [Reptiles of the plains of Turkmenistan, in Russian]. Instit. Zool., ANTur SSR, Ashkhabad 308 pp.
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, Roberto; Alberto Venchi, Giuseppe M. Carpaneto, Marco A. Bologna 2000. The Reptiles of Anatolia: a Checklist and Zoogeographical analysis. Biogeographia, 21-2000: 441-554. - get paper here
  • Siroky, P.; Stuchlik, S.; Fritz, U. & Moravec, J. 2009. Basic morphological data of native Czech Emys orbicularis revealed by subfossil finds. Biologia 64/4: 795—797 - get paper here
  • Sobolenko, L.Yu., Nekrasova, O.D., Sorokina, S.I., Moroz, L.M. 2018. Current state of reptiles of western podillia. Ukrainian Journal of Ecology, 8(1): 706–711 - get paper here
  • Sommer, Robert S.; Arne Persson, Nina Wieseke & Uwe Fritz 2007. Holocene recolonization and extinction of the pond turtle, Emys orbicularis (L., 1758), in Europe. Quaternary Science Reviews 26: 3099–3107 - get paper here
  • Sos, T. 2008. Review of recent taxonomic and nomenclatural changes in European Amphibia and Reptilia related to Romanian herpetofauna. Herpetologica Romanica 2: 61-91
  • Sos, Tibor S 2013. Conservation activities for European pond turtles (Emys orbicularis) in Romania. Herpetology Notes 6: 147-148
  • Stoev, Pavel 2000. On the distribution, biology and ecology of amphibians and reptiles in the Derventski Heights and the Sakar Mountain, South-East Bulgaria Hist. Nat. Bulgarica 12: 59–69 - get paper here
  • Strugariu, A., Zamfirescu, A.R., Nicoară, A., Gherghel, I., Sas, I., Puşcaşu, C.M. & Bugeac, T. 2008. Preliminary data regarding the distribution and status of the herpetofauna in Iaşi County (Romania). North-Western Journal of Zoology 4 (Suppl.1): S1-S23 - get paper here
  • Stuckas, Heiko; Guillermo Velo-Antón, Soumia Fahd, Mohsen Kalboussi, Rachid Rouag, Marco Arculeo, Federico Marrone, Francesco Sacco, Melita Vamberger, Uwe Fritz 2014. Where are you from, stranger? The enigmatic biogeography of North African pond turtles (Emys orbicularis). Organisms Diversity & Evolution, DOI:<br />10.1007/s13127-014-0168-4 - get paper here
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp. - get paper here
  • Thienpont, Stéphanie; Jean-Pierre Vacher, Maud Berroneau, André Miquet, Laurent Barthe, Aurélie Caillebotte, Damien Lerat , Thomas Gendre, Zoey Owen-Jones, Jean-Luc Marandon, Raphaël Quesada. 2013. Conservation activities for European pond turtles (Emys orbicularis) in France. Herpetology Notes 6: 139-141
  • Tóth, Tamás, László Krecsák, Thomas Madsen and Beáta Újvári 2002. Herpetofaunal locality records on the Greek Islands of Corfu (Amphibia, Reptilia). Herpetozoa 15 (3/4):149-169 - get paper here
  • Toth, Tamas; Miklós Heltai, Andrea Keszi, Goran Susic, Levente Moharos, Balazs Farkas, Csaba Geczy, Orsolya Torda, Janos Gal 2017. Herpetofauna inventory of the small islands of the Cres-Losinj Archipelago (North Adriatic Sea, Croatia) (Amphibia; Reptilia). Herpetozoa 30 (1/2): 21 - 28 - get paper here
  • Trakimas, Giedrius; Jonas Sidaravičius 2008. Road mortality threatens small northern populations of the European pond turtle, Emys orbicularis. Acta Herpetologica 3 (2): 161-166
  • Trapp, B. 2007. Reptilien und Amphibien des griechischen Festlandes. Natur und Tier Verlag (Münster), 300 pp. - get paper here
  • Trutnau, L. 1975. Europaische Amphibien und Reptilien. Belser Verlag, Stuttgart 212 pp.
  • TTWG; Anders G.J. Rhodin, John B. iverson, Roger Bour, Uwe Fritz, Arthur GeoRges, H. BRadley Shaffer, and Peter Paul van Dijk 2017. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (8th Ed.). In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Pritchard, P.C.H., and Mittermeier, R.A. (Ed Chelonian Research Monographs 7:1–292. doi: 10.3854/crm.7.checklist.atlas.v8.2017 - get paper here
  • TTWG; Rhodin, A.G.J.; van Dijk, P.P.; Iverson, J.B. & Shaffer, H.B. [turtle taxonomy working group] 2010. Turtles of the World, 2010 Update: Annotated Checklist of Taxonomy, Synonymy, Distribution, and Conservation Status. Chelonian Research Monographs (ISSN 1088-7105) No. 5, doi:10.3854/crm.5.000.checklist.v3.2010 - get paper here
  • Uhrin, Marcel; Peter Havaš, Martin Minařík, Karel Kodejš, Imrich Bugoš, Stanislav Danko, Tomáš Husák, Daniel Koleska and Daniel Jablonski 2016. Distribution updates to amphibian and reptile fauna for the Republic of Macedonia. Herpetology Notes 9: 201-220 - get paper here
  • Valenzuela, Alejandro; Miguel Ángel Cau and Josep Antoni Alcover 2016. Archaeological evidence for the introduction of Emys orbicularis (Testudines: Emydidae) in the Balearic Islands, Western Mediterranean. Amphibia-Reptilia 37 (2): 229-236 - get paper here
  • Vamberger, M., & Fritz, U. 2018. Big data can cause big mistakes: using the Societas Europaea Herpetologica atlas by Sillero et al. (2014), the distribution of Emys orbicularis will be misunderstood. Biologia - get paper here
  • VAMBERGER, M., LIPOVŠEK, G., ŠALAMUN, A., CIPOT, M., FRITZ, U. & GOVEDIČ, M. 2017. Distribution and population size of the European pond turtle Emys orbicularis in Ljubljansko barje, Slovenia. Vertebrate Zoology 67 (2): - get paper here
  • Vamberger, M., Stuckas, H., Sacco, F., D'Angelo, S., Arculeo, M., Cheylan, M., Corti, C., Lo Valvo, M., Marrone, F., Wink, M., Fritz, U. 2015. Differences in gene flow in a twofold secondary contact zone of pond turtles in southern Italy (Testudines: Emydidae: Emys orbicularis galloitalica, E. o. hellenica, E. trinacris). Zoologica Scripta 44: 233–249 - get paper here
  • Velo-Antón, Guillermo, Paulo Pereira, Soumia Fahd, José Teixeira and Uwe Fritz. 2015. Out of Africa: did Emys orbicularis occidentalis cross the Strait pf Gibraltar twice? Amphibia-Reptilia 36 (2): 133-140 - get paper here
  • Velo-Antón, Guillermo. 2013. Towards better management of European pond turtles (Emys orbicularis) in conservation programs. Herpetology Notes 6: 381-382.
  • Velo-Antón, Guillermo; García-París, Mario; Cordero-Rivera, Adolfo 2008. Patterns of nuclear and mitochondrial DNA variation in Iberian populations of Emys orbicularis (Emydidae): conservation implications. Conservation Genetics. 9:1263–1274 - get paper here
  • Velo-Antón, Guillermo; Bahija el Marnisi, Uwe Fritz & Soumia Fahd 2015. Distribution and conservation status of Emys orbicularis in Morocco. Vert. Zool. 65(1):131–134
  • Vignoli, Leonardo, Marco A. Bologna, Silvia Manzini, Lorenzo Rugiero and Luca Luiselli. 2015. Attributes of basking sites of the European Pond Turtle (Emys orbicularis) in central Italy. Amphibia-Reptilia 36 (2): 125-131 - get paper here
  • Webb, R. G. 1993. Emys Dumeril, 1806 (Reptilia, Testudines): proposed conservation. Bull. Zool. Nomenclature 50 (3): 224-227 - get paper here
  • Williams, E. E. 1959. Cervical ribs in turtles. Breviora (101): 1-13 - get paper here
  • Wolff, Bernd 2015. Langjährige Erfahrungen bei der Haltung und Nachzucht der Europäischen Sumpfschildkröte. Teil 1: Haltung, Gestaltung einer Freilandanlage, Fütterung. Terraria-Elaphe 2015 (3): 24-31 - get paper here
  • Wolff, Bernd 2015. Langjährige Erfahrungen bei der Haltung und Nachzucht der Europäischen Sumpfschildkröte. Teil 2: Eiablage, Inkubation, Aufzucht der Jungtiere, Überwinterung, Haltungsprobleme. Terraria-Elaphe 2015 (4): 62-70 - get paper here
  • Wütschert, R. 1984. Neues uber die Reptilienfauna der Insel Korfu. Salamandra 20 (4): 221-228 - get paper here
  • Ziane, Nadia; Samira Fediras, rachid Rouag, Anthony Olivier, slim Benyacoub 2020. Feeding habits of the European pond turtle (Emys orbicularis) in the Lake Tonga, Algeria. Herpetology Notes 13: 875-881 - get paper here
  • Zuffi, Marco A.L. and Elena Foschi. 2015. Reproductive patterns of European pond turtles differ between sites: a small scale scenario. Amphibia-Reptilia 36 (4): 339-349 - get paper here
  • Zuffi, Marco A.L.; Simona Citi, Elena Foschi, Francesca Marsiglia, Eva Martelli 2015. Into a box interiors: clutch size variation and resource allocation in the European pond turtle. Acta Herpetologica 10: 39-45 - get paper here
  • Кhahibullin, V.F. 2000. DISTRIBUTION OF EUROPEAN POND TURTLE EMYS ORBICULARIS (L., 1758) IN BASHКIRIA (SOUTH URALS, RUSSIA). Biodiversity and Dynamics of Ecosystems in North Eurasia, Vol. 3. DIVERSITY OF ТНЕ FAUNA OF NORTH EURASIA, Part 1: 159-160
External links  
Is it interesting? Share with others:

As link to this species use URL address:

without field 'search_param'. Field 'search_param' is used for browsing search result.

Please submit feedback about this entry to the curator