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Eutropis borealis (BROWN & ALCALA, 1980)

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Higher TaxaScincidae, Mabuyinae, Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymMabuya multicarinata borealis BROWN & ALCALA 1980
Mabuya multicarinata borealis — OTA 1991
Mabuya multicarinata borealis — LAGAT 2009
Eutropis multicarinata borealis — Mausfeld and Schmitz 2003: 169
Eutropis multicarinata borealis — GAULKE 2011
Eutropis multicarinata borealis — BROWN et al. 2013
Eutropis borealis — BARLEY et al. 2020: 59 
DistributionPhilippines (Catanduanes, Luzon, Polillo, Babuyan Claro, Batanes)

Type locality: Subic Bay area, Luzon Island, Philippines.  
Reproduction 
TypesHolotype: CAS 15447, collected by J. Thomson, 7 June 1907.
Paratypes: CAS, BMNH, FMNH
Additional specimens: KU 
DiagnosisDiagnosis: A species of Eutropis, distinguished by the following combination of characters: (1) body medium-sized (SVL 64–83 mm in adults); (2) interparietal small, parietals in contact posteriorly; (3) paravertebrals 37–42; (4) sum of subdigital lamellae on all five toes of one foot 80–89; (5) ventral scales rows 25–30; (6) midbody scale rows 28–33; (7) keels on the dorsal and lateral body scales moderately defined, 6–11; (8) lower eyelid scaly; (9) supraciliaries five; (10) prefrontals separated; (11) primary temporal scales one; (12) dorsal and lateral body surfaces with relatively uniform bronze and dark brown coloration, respectively, without pronounced light stripes.

Comparisons: Critical comparisons for E. borealis include other Philippine species of Eutropis, particularly those known from the northern islands of the archipelago. Eutropis borealis can be distinguished from members of the E. indeprensa complex (including both E. lapulapu and E. cumingi, with which it occurs sympatrically) by its larger (adult SVL 64–83 mm), more robust body (vs. 45–70 mm in E. lapulapu and 43–60 mm in E. cumingi) and fewer subdigital toe lamellae (80–89 in E. borealis vs. 70–80 in E. lapulapu or 59–70 in E. cumingi). Eutropis borealis can also be distinguished from E. cumingi by fewer vertebral scale rows (37–42 vs. 42–47). It can be distinguished from E. multifasciata by its smaller body size (64–83 mm vs. 101–141 mm adult SVL), and more numerous and pronounced (Fig. 8) keels on the trunk dorsals (6–9 vs. 3). Eutropis borealis can be readily distinguished from E. bontocensis by color pattern, which either lacks lateral stripes or has faint, lateral, light stripes on the anterior portion of the body (compared to two prominent, light stripes on the lateral surface that extend the length of the body). It can also be distinguished from E. bontocensis by more strongly keeled dorsal scales (vs. lightly keeled dorsal scales), by a larger number of subdigital toe lamellae (80–89 vs. 67–76), and fewer vertebral scale rows (37–42 vs. 44–50). A phenotypically very similar species (E. gubataas) appears to be patchily distributed across Northern Luzon Island and the Babuyan Island Group. It can readily be distinguished from E. borealis using genetic data (Fig. 5).

Coloration in life: Dorsal ground coloration and tail an iridescent bronze to olive coloration, usually with scattered dark brown flecks; head and neck brown. Lateral surfaces with a thick, dark brown band that extends from the eye, past the hindlimb, becoming thinner and less distinct posteriorly. Chin creamy white with dark markings. Dorsal surfaces of forelimbs, hindlimbs, and digits dark with indistinct spots. Frequently there are traces of light lines above and below the lateral brown stripes. 
CommentDistribution: possibly many other islands, but their identity is not clear (Barley et al. 2020: 60). 
References
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