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Eutropis resetarii BATUWITA, UDUGAMPALA & EDIRISINGHE, 2020

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Higher TaxaScincidae, Mabuyinae, Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymEutropis resetarii BATUWITA, UDUGAMPALA & EDIRISINGHE 2020
Eutropis beddomii — SMITH 1935 (in part)
Eutropis beddomii — DERANIYAGALA 1953 (in part)
Eutropis carinata — BATUWITA 2016 (in part) 
DistributionSri Lanka

Type locality: Agra Arboretum, near Torrington Estate, Agarapatana, Sri Lanka, 06°51’N, 80°41’E, elevation 1550 m a.s.l.  
Reproduction 
TypesHolotype: WHT 6759, adult male, SVL 121.0 mm, 26 December 2003, collected by Sudesh Batuwita and Kalana Prasad Maduwage.
Paratypes: Three paratypes: WHT 6771, adult male, SVL 119.0 mm, same location data as holotype, 24 December 2003, collected by Mohomed Mujthaba Bahir, Anjana Silva and Sudesh Batuwita; WHT 6772, adult female, SVL 112.0 mm, same location data as holotype, 12 May 2004; WHT 6773, subadult male, SVL 74.0 mm, same location data as holotype, 28 December 2003, collected by Mohomed Mujthaba Bahir and Sudesh Batuwita. 
DiagnosisDiagnosis: Distinguished from all other species of Eutropis by the combination of the following characters: supranasals narrowly separated (not touching); prefrontals widely separated; frontal twice as long as wide; only the second supraocular in contact with frontal; frontoparietals paired, as wide as long, in contact with second, third and fourth supraoculars; the medial border of the fourth supraocular completely in contact with frontoparietal; parietals completely separated by interparietal; one pair of nuchals; 6 or 7 supraciliaries; first three pairs of supraciliaries in contact with the first supraocular; two pretemporals, both in contact with parietal; two primary temporals; two secondary temporals, separated; seven supralabials; two post-supralabials; postmental completely in contact with the first and the second infralabials; three pairs of enlarged chin shields, first pair separated by a median scale, second pair separated by a single scale and the third pair separated by three scales; third pair of chin shields not contacting/narrowly contacting second pair of chin shields; third pair also separated from the infralabial row by sublabial scale row; 40–45 paravertebrals; 55–57 ventrals; 30 transverse scale rows across midbody; subdigital lamellae under 4th digit of manus, 11–12 and pes, 14–15; dark brown dorsal coloration; and having five dark longitudinal stripes on dorsum (excluding the line confluent with dorsolateral stripe on each side).
Eutropis resetarii sp. nov. can be distinguished from Eutropis carinata by the following suite of characters (Figs. 2–5): first loreal does not reach the dorsal surface of snout (vs. reaches in E. carinata); lower preocular larger (vs. smaller) than the anterior loreal scale; first three (vs. first two and partially the third) supraciliaries completely touching the first supraocular; third pair of chin shields not in contact or in narrow (vs. broad) contact with second pair of chin shields; 30 (vs. 32) transverse scale rows across midbody; digits comparatively robust (vs. slender); olive brown (vs. dark copper brown) dorsal coloration; having granular (vs. pointed) ear lobules; and having a greater external ear opening size.
Eutropis resetarii sp. nov. differs from Eutropis lankae in that the first loreal does not reach the dorsal surface of snout (vs. reaches in E. lankae); lower preocular larger (vs. smaller) than the anterior loreal scale; first three (vs. first two and partially the third) supraciliaries completely touching the first supraocular; lateral border of postmental completely in contact with the first and the second (vs. first and partially the second) infralabials; third pair of chin shields not in contact or in narrow (vs. broad) contact with second pair of chin shields (Figs. 3A vs. 3B; 3C vs. 3D; 3E vs. 3F); palm and sole scales rounded, more or less juxtaposed (vs. tubercle-like imbricate scales); digits comparatively robust (vs. slender); olive brown (vs. dark copper brown) dorsal coloration; having granular (vs. pointed) ear lobules; and having a greater external ear opening size, 40– 46% (vs. 23–38%) of eye diameter.primary temporals; two secondary temporals, separated; seven supralabials; two post-supralabials; postmental completely in contact with the first and the second infralabials; three pairs of enlarged chin shields, first pair separated by a median scale, second pair separated by a single scale and the third pair separated by three scales; third pair of chin shields not contacting/narrowly contacting second pair of chin shields; third pair also separated from the infralabial row by sublabial scale row; 40–45 paravertebrals; 55–57 ventrals; 30 transverse scale rows across midbody; subdigital lamellae under 4th digit of manus, 11–12 and pes, 14–15; dark brown dorsal coloration; and having five dark longitudinal stripes on dorsum (excluding the line confluent with dorsolateral stripe on each side).
Eutropis resetarii sp. nov. can be distinguished from Eutropis carinata by the following suite of characters (Figs. 2–5): first loreal does not reach the dorsal surface of snout (vs. reaches in E. carinata); lower preocular larger (vs. smaller) than the anterior loreal scale; first three (vs. first two and partially the third) supraciliaries completely touching the first supraocular; third pair of chin shields not in contact or in narrow (vs. broad) contact with second pair of chin shields; 30 (vs. 32) transverse scale rows across midbody; digits comparatively robust (vs. slender); olive brown (vs. dark copper brown) dorsal coloration; having granular (vs. pointed) ear lobules; and having a greater external ear opening size.
Eutropis resetarii sp. nov. differs from Eutropis lankae in that the first loreal does not reach the dorsal surface of snout (vs. reaches in E. lankae); lower preocular larger (vs. smaller) than the anterior loreal scale; first three (vs. first two and partially the third) supraciliaries completely touching the first supraocular; lateral border of postmental completely in contact with the first and the second (vs. first and partially the second) infralabials; third pair of chin shields not in contact or in narrow (vs. broad) contact with second pair of chin shields (Figs. 3A vs. 3B; 3C vs. 3D; 3E vs. 3F); palm and sole scales rounded, more or less juxtaposed (vs. tubercle-like imbricate scales); digits comparatively robust (vs. slender); olive brown (vs. dark copper brown) dorsal coloration; having granular (vs. pointed) ear lobules; and having a greater external ear opening size, 40– 46% (vs. 23–38%) of eye diameter (Batuwita et al. 2020).

Color in life: Sexes alike during non-breeding period. Olive brown dorsum with longitudinally arranged black spots forming an uninterrupted series of five longitudinal lines, another two lines on each side begin just before the forelimb origin; half a scale-width to one scale-width distinct yellowish-brown dorsolateral stripe begins from supraciliaries to body and on to tail-base; lateral sides of body (from behind ear opening) more or less bicolored, upper half dark brown with scattered light brown spots; lower half light brown and each scale with dark anterior marking; ventral side of breeding males bright yellow; females with dusky white belly; limbs brown. Juvenile color copper brown with yellowish-brown lateral stripe; lateral sides dark brown; and limbs dark (Fig. 5A in Batuwita et al. 2020). 
Comment 
EtymologyThe species name is a patronym in the Latin genitive singular, in honour of Alan Resetar (Collections Manager of the Division of Amphibians and Reptiles at the Field Museum of Natural History, Chicago, USA), for his contributions to the conservation of herpetofauna. 
References
  • Batuwita S, Udugampala S, Edirisinghe U. 2020. Description of a new species of Eutropis (Sauria: Scincidae) from the Central Hills of Sri Lanka with the resurrection of Eutropis lankae (Deraniyagala). JAD 2020; 2 (2) - get paper here
  • Batuwita, Sudesh 2016. Description of Two New Species of Eutropis (Reptilia: Scincidae) from Sri Lanka with a Redescription of Eutropis madaraszi (Méhely). Journal of Herpetology 50 (3): 486-496. - get paper here
  • Deraniyagala, P.E.P. 1953. A coloured atlas of some vertebrates from Ceylon. Vol. 2. Tetrapod Reptilia. Govt. Press, Colombo, 101 pp.
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
 
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