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Gehyra ipsa HORNER, 2005

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Bungle Bungle Ranges Gehyra 
SynonymGehyra koira ipsa HORNER 2005
Gehyra koira ipsa — OLIVER et al. 2010
Gehyra koira ipsa — WILSON & SWAN 2013: 122
Gehyra ipsa — COGGER 2014: 359
Gehyra koira ipsa — ELLIS et al. 2018
Gehyra ipsa — OLIVER et al. 2019
Gehyra ipsa — OLIVER et la. 2020: 40 
DistributionAustralia (Western Australia)

Type locality: Piccaninny Massif, Purnululu (Bungle Bungle)
National Park, Western Australia, 17°27’S 128°24’E.  
TypesHolotype: WAM R.101238, adult male; paratypes: NTM, WAM 
DiagnosisDiagnosis. A large Gehyra species (up to 94.9 mm SVL), differing from all other Gehyra species outside of the G. koira complex as per the diagnosis above. Differs from other members of the G. koira complex in the combination of: large size within complex (adult SVL up to 94.9 mm, mean 84.9 mm, largest of the G. koira complex); pores in males moderately numerous (15–18); first chin shield pair bordered posteriorly by a single enlarged medial scale, ∼1.5–4 times the size of adjacent scales and always the largest in first row of gular scales posterior to chin shields; second chin shields approximately two thirds length of first chin shields (mean ratio 0.78, range 0.75–0.81); adults in base colouration with tan to brownish dorsum with light transverse barring, a distinct to indistinct brown postorbital stripe, tails usually with at least some distinct transverse light and/or dark barring.
Further diagnosed from other species within the G. koira complex genetically by four unique amino acids in the ND2 locus (Table 1).
Gehyra ipsa is likely to occur in parapatry or even sympatry with G. koira (which it is morphologically most similar to), G. gemina sp. nov. and possibly G. calcitectus sp. nov. It can often be differentiated from all three by the combination of larger size (mean
and maximum adult SVL, respectively: 84.9 mm and 94.9 mm), presence of an enlarged medial scale behind the first pair of chin shields, which is always the largest in the first row of gular scales posterior to chin shields (vs. absence or, if present, not the largest scales in first gular row) and dorsal pattern comprising transverse light bands (vs. often comprising light pale tan ocelli or blotches in G. calcitectus sp. nov.). 
CommentSimilar species: Due to the morphological overlap and potential occurrence of sympatry of G. koira and G. ipsa, in addition to G. calcitectus sp. nov., there remains some uncertainty as to which species some specimens apply. 
EtymologyNamed after the Greek noun koira, meaning king, ruler or commander, in reference to Max King, in recognition of his landmark work on the cytology and taxonomy of Gehyra. The name is intended as a noun in apposition. The subspecifi c epithet is from the Latin word ipsa, meaning to make prominent one of two or more subjects, in reference to the morphological divergence of the Purnululu population from conspecifi c populations. The name is here intended as a noun in apposition. 
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Horner, P. 2005. Gehyra koira sp. nov. (Reptilia: Gekkonidae), a new species of lizard with two allopatric subspecies from the Ord-Victoria region of north-western Australia and a key to the Gehyra australis species complex. The Beagle 21: 165-174
  • Oliver PM, Prasetya AM, Tedeschi LG, Fenker J, Ellis RJ, Doughty P, Moritz C. 2020. Crypsis and convergence: integrative taxonomic revision of the Gehyra australis group (Squamata: Gekkonidae) from northern Australia. PeerJ 8:e7971 - get paper here
  • Oliver, P. M., Ashman, L. G., Bank, S., Laver, R. J., Pratt, R. C., Tedeschi, L. G., & Moritz, C. C. 2019. On and off the rocks: persistence and ecological diversification in a tropical Australian lizard radiation. BMC Evolutionary Biology, 19(1), 81 - get paper here
  • OLIVER, PAUL; MARK SISTROM, BURHAN TJATURADI, KELIOPAS KREY & STEPHEN RICHARDS 2010. On the status and relationships of the gecko species Gehyra barea Kopstein 1926, with description of new specimens and a range extension. Zootaxa 2354: 45–55 - get paper here
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
  • Wilson, S. & Swan, G. 2013. A complete guide to reptiles of Australia, 4th ed. New Holland Publishers, 592 pp.
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