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Gloydius intermedius (STRAUCH, 1868)

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Central Asian pitviper
Russian: Средний щитомордник
Chinese: 中介蝮 
SynonymTrigonocephalus intermedius STRAUCH 1868: 295
Agkistrodon halys intermedius STRAUCH 1876
Halys intermedia — PETERS 1877: 735
Ancistrodon intermedius — BOULENGER 1890: 140
Ancistrodon intermedius — BOULENGER 1896: 525
Agkistrodon blomhoffi intermedius — STEJNEGER 1907: 464
Ancistrodon blomhoffi intermedius — DESPAX 1913: 181
Ancistrodon halys intermedius — EMELIANOV 1929: 130
Ancistrodon halys intermedius — RENDAHL 1933: 15
Ancistrodon saxatilis EMELIANOV 1937
Agkistrodon saxatilis GLOYD 1972: 569
Agkistrodon halys cognatus GLOYD 1977
Agkistrodon halys intermedius — HARDING & WELCH 1980
Agkistrodon shedaoensis continentalis ZHAO 1980: 12
Agkistrodon saxatilis — SCHNEEWEISS 1989
Agkistrodon intermedius saxatilis — WELCH 1994: 13
Agkistrodon intermedius intermedius — WELCH 1994: 11
Agkistrodon intermedius — ORLOV & BARABANOV 1999
Gloydius saxatilis — MCDIARMID, CAMPBELL & TOURÉ 1999: 307
Gloydius intermedius — MCDIARMID, CAMPBELL & TOURÉ 1999: 306
Gloydius intermedius — GUMPRECHT et al. 2004
Gloydius saxatilis — JIANG & ZHAO 2009
Gloydius intermedius — WALLACH et al. 2014: 311
Gloydius saxatilis — WALLACH et al. 2014: 311
Gloydius intermedius — DAVID & VOGEL 2015
Gloydius intermedius — WAGNER et al. 2015
Gloydius saxatilis — PARK et al 2022 
DistributionN China (incl. Liaoning, Jilin, Heilongjiang),
E Russia, North Korea, South Korea

Type locality: Irkutsk, Eastern Siberia; Terra typica restricta (hoc loco). Malyi Hinggan Mountain Range (Xiao Hinggan Ling), Cape Tyr on the Amur River and Cape Kofel on the Ussuri River, Russian Far East Region.

saxatilis: NE China south to Shandong,
Russia (east of Selemdzha River in E Siberia, Primorskye Kray and Khabarovsk Kray, Far East), Korea; Type locality: Vladivostok River, Russia  
Reproductionovovivparous 
TypesLectotype. ZISP 2221. Prom. Tyr (Amur). Leg.: Schrenck, 1855 
DiagnosisDiagnosis. Total length up to 800 mm and more. Dorsal scales in 23 (rarely 21) rows at midbody. Supralabials 7 – 8; ventrals 148 – 175; subcaudals 34–52. 
CommentVenomous!

Synonymy partly after KHALIKOV & ANANJEVA (pers. comm.). G. saxatilis is not recognized by GUMPRECHT et al. 2004. ORLOV & BARABANOV 1999 synonymized G. saxatilis with G. intermedius. See Jiang & Zhao 2009 for comparisons of G. saxatilis, shedaoensis, and their new species lijianlii (which has been synonymized with G. changdaoensis).

Distribution: see map in ORLOV & BARABANOV 1999.

NCBI taxonID: 92067 [saxatilis] 
References
  • Adnagulov E.V. 2016. On the distribution of amphibians and reptiles in the Evreiskaya Autonomous Oblast' (Russia). [in Russian] Current Studies in Herpetology, 16(3/4): 87–106 - get paper here
  • Boulenger, G.A. 1896. Catalogue of the snakes in the British Museum, Vol. 3. London (Taylor & Francis), xiv + 727 pp. - get paper here
  • Boulenger, George A. 1890. A list of the reptiles and batrachians of Amoorland [Russia]. Ann. Mag. nat. Hist. (6) 5: 137-144 - get paper here
  • Chen, Y.; Zhang, D.; Jiang, K. & Wang, Z. 1992. Evaluation of Snake Venoms among Agkistrodon Species in China. Asiatic Herpetological Research 4: 58-61 - get paper here
  • David, P. & Vogel, G. 2015. An updated list of Asian pitvipers and a selection of recent publications, in: Visser: Asian Pitvipers. Breeding Experience & Wildlife. Edition Chimaira, Frankfurt am Main, 571pp. ISBN 978-3-89973-450-8 - get paper here
  • Despax, RAYMOND 1913. Sur une collection de reptiles et de batraciens rassemblée par M. le Dr. Legendre dans les marches Thibétaines. Bull. Mus. Hist. Nat., Paris, [ser. 1], 19 (4): 179-183 - get paper here
  • Do, Min Seock; Ki-Baek Nam 2021. Distribution Patterns and Ecological Niches of the Red-tongued Pit Viper (Gloydius ussuriensis) and the Central Asian Pit Viper (Gloydius intermedius) in Cheonmasan Mountain, South Korea. Russian Journal of Herpetology 28 (6): 348-354
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  • Koo, Kyo Soung 2019. First Report on the Hybridization between Gloydius brevicaudus and G. intermedius. Herpetology Notes 12: - get paper here
  • Li, J.L. 1999. Infraspecific classification of two species of Gloydius (Serpentes: Crotalinae). Acta Zootaxonomica Sinica 24(4), Oct 1999:454-460
  • LIN Xuan-Long, GUO Dong-Sheng 2012. A New Record of Snake in Beijing: Gloydius intermedius. Chinese Journal of Zoology 2012, 47(2):136-137 - get paper here
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  • Orlov, Nikolai L. and Andrei V. Barabanov. 1999. Analysis of nomenclature, classification, and distribution of the Agkistrodon halys - Agkistrodon intermedius complexes: a critical review. Russ. J. Herpetol. 6 (3): 167-192 - get paper here
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  • Peters, W. 1877. Eine Übersicht über die während der sibirischen Expedition von 1876 von Hrn. Dr. O. Finsch gesammelten Säugethiere, Amphibien und Fische. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin 1877: 734-738 - get paper here
  • Rendahl, Hialmar 1933. Die Unterarten des Ancistrodon halys Pall. nebst elnigen Bemerkungen zur Herpetologie Zentralasiens. Arkiv för Zoologi 25 (8): 1-31
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