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Goggia rupicola (FITZSIMONS, 1938)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Namaqua Dwarf Leaf-toed Gecko 
SynonymPhyllodactylus lineatus rupicolus FITZSIMONS 1938: 162
Phyllodactylus lineatus rupicolus — KLUGE 1993
Phyllodactylus rupicolus — BRANCH et al. 1995: 40
Goggia rupicola — BAUER et al. 1997
Goggia rupicola — RÖSLER 2000: 83
Goggia rupicola — BATES et al. 2014: 113 
DistributionRepublic of South Africa;

Type locality: between O´okiep and Springbok, Little-Namaqualand  
Reproductionoviparous 
TypesHolotype: DNMNH (= TM) 18161 
DiagnosisDiagnosis: Distinguished from other members of the P. lineatus complex by a combination of the following features: usually four preanal pores in males, a high number of midbody scale rows (80-90), 10-12 scales between the nostril and anterior margin of the orbit, usually 2 (1-3) nasorostrals. and a dorso-ventrally flattened head and body. In addition, it differs by 4-6 fixed alleles from the other taxa examined electrophoretically (Branch et al. 1995: 40).

Description: Head dorso-ventrally flattened (depth 2.03 times width; range 1.63-2.57); snout rounded, and at least 1.5 times the eye diameter; ear opening small, obliquely rounded, and without enlarged lobules or a tympanic shield; rostral subpentagonal with a median cleft above; nostril pierced between rostra!, 1st supralabial and three nasals, the largest bordering the rostral; usually 2 (50%) nasorostral granules, rarely 1 (23.3%) or 3 (26.7%); supralabials usually 7 (50%), often 6 (25%), less often 8 (21.4%) or 9 (3.6%); infralabials usually 7 (50%), often 8 (42.5%), rarely 6 (7.5%); mental subpentagular bordered by 2-3 (average 2.9) chinshields that are larger than adjacent granules; 5-7 (average 6.28) granules bordering chinshields. Body short (maximum snout-vent length 31.51 mm) and dorsoventrally flattened, covered dorsally with uniform. smooth, flattened, subimbricate granules; belly covered in smooth. hexagonal, imbricate scales that are much larger than on back. Limbs moderate and covered in uniform, flattened, subimbricate granules; digits flattened basally, with three rows of subdigital granules, the median series slightly broader; toe tips rounded, only slightly expanded and bearing a pair of large, subrectangular scansors that enclose a small claw. An angular series of 4 (80%, n = 8), rarely 3 (20%) preanal pores present only in males; two to three enlarged tubercnlar scales on either side of tail base near vent, that are larger in males. Original tail cylindrical, tapering, and up to 1.21 times snout-vent length; covered above with regular rows of uniform, smooth grannles, that are much larger, flattened and imbricate on ventral surface (Branch et al. 1995: 40, Figure 1D). 
Comment 
EtymologyRupicolous means “inhabiting rocks and stones”. 
References
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bauer A M. Good D A. Branch W R. 1997. The taxonomy of the Southern African leaf-toed geckos (Squamata: Gekkonidae), with a review of Old World "Phyllodactylus" and the description of five new genera. Proc. Cal. Acad. Sci. 49 (14): 447-497. - get paper here
  • Branch, W. R., A. M. BAUER & D. A. GOOD 1995. Species limits in the Phyllodactylus lineatus complex (Reptilia: Gekkonidae), with the elevation of two taxa to specific status and the description of two new species. J. Herp. Assoc. Africa 44 (2): 33-54. - get paper here
  • Fitzsimons,V.F.M. 1938. Transvaal Museum Expedition to South-West Africa and Little Namaqualand, May to August 1937 - Reptiles and Amphibians. Annals Transvaal Mus. 19 (2): 153-209 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
 
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