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Gonatodes albogularis (DUMÉRIL & BIBRON, 1836)

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Higher TaxaSphaerodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
SubspeciesGonatodes albogularis albogularis (DUMÉRIL & BIBRON 1836)
Gonatodes albogularis bodinii RIVERO BLANCO 1964
Gonatodes albogularis fuscus (HALLOWELL 1855)
Gonatodes albogularis notatus (REINHARDT & LÜTKEN 1862) 
Common NamesE: White-throated Clawed Gecko, Yellow-headed Gecko, White-throated Gecko
G: Gelbkopf-Gecko, Weisskehl-Gecko
S: Limpiacasa 
SynonymGymnodactylus albogularis DUMÉRIL & BIBRON 1836: 415
Gymnodactylus maculatus STEINDACHNER 1869: 16 (fide BOULENGER 1885)
Gonatodes albogularis — BOULENGER 1885: 59
Goniodactylus albogularis — COPE 1885: 182
Gonatodes albigularis — HEYER 1967 (in error)
Gonatodes albogularis — CONANT & COLLINS 1991: 87
Gonatodes albogularis — SCHWARTZ & HENDERSON 1991: 408
Gonatodes albogularis — LINER 1994
Gonatodes albogularis — KÖHLER 2000: 48
Gonatodes albogularis — MCNISH 2011
Gonatodes albogularis — MCCRANIE 2018
Gonatodes albogularis — MARTÍNEZ-FONSECA et al. 2024

Gonatodes albogularis bodinii RIVERO BLANCO 1964
Gonatodes bodinii RIVERO BLANCO 1964
Gonatodes bodinii — DONOSO-BARROS 1968: 108
Gonatodes bodinii — PETERS et al. 1970: 132
Gonatodes albogularis bodinii — RÖLL 2009

Gonatodes albogularis fuscus (HALLOWELL 1855)
Stenodactylus fuscus HALLOWELL 1855: 33
Gymnodactylus fuscus — DUMÉRIL 1856: 477
Gonatodes albogularis fuscus — BOULENGER 1885: 59
Gonatodes fuscus — STEJNEGER 1917: 264
Gonatodes fuscus — CARR 1939
Gonatodes fuscus — GRANT 1940
Gonatodes fuscus — BURT & MYERS 1942
Gonatodes fuscus — SMITH 1944
Gonatodes fuscus — SMITH 1946
Gonatodes fuscus — MERTENS 1952: 38
Gonatodes fuscus — TAYLOR 1956: 27
Gonatodes albogularis fuscus — VANZOLINI & WILLIAMS 1962: 491
Gonatodes fuscus — STUART 1963
Gonatodes albogularis fuscus — WERMUTH 1965: 42
Gonatodes fuscus — NEMURAS 1966
Gonatodes fuscus — RUIBAL 1968
Gonatodes albogularis fuscus — SCHWARTZ & HENDERSON 1988: 130
Gonatodes fuscus — RÖLL 2009
Gonatodes albogularis fuscus — POWELL & HENDERSON 2012

Gonatodes albogularis notatus (REINHARDT & LÜTKEN 1862)
Gymnodactylus notatus REINHARDT & LÜTKEN 1862: 280
Gonatodes notatus — BARBOUR 1937: 108
Gonatodes notatus — COCHRAN 1941
Gonatodes albogularis notatus — VANZOLINI & WILLIAMS 1962: 492
Gonatodes albogularis notatus — WERMUTH 1965: 42
Gonatodes albogularis notatus — SCHWARTZ & HENDERSON 1988
Gonatodes albogularis notatus — POWELL & HENDERSON 2012: 127 
DistributionUSA (introduced in Florida)
Antilles, Martinique, Cuba, Hispaniola, Jamaica, Haiti (Acquin), Grand Cayman, Cuba
Mexico (Chiapas, Veracruz), El Salvador, Costa Rica, Nicaragua, Panama, Guatemala, Belize, Honduras,
Colombia (Valle del Cauca, Chocó, Putumayo, Caquetá, Meta, Arauca, Guainia, Vaupés), Venezuela (Barinas etc.)

bodinii: Venezuela (Monjes Islands: Monje Grande del Sur, Monje Grande del Norte); Type locality: Monje Grande del Sur, Venezuela.

fuscus: Costa Rica, Panama; Cuba (incl. Cayo Santa María);
USA: introduced in southern Florida (Schmidt 1953);
Type locality: “Nicaragua”; restricted to Rama, Nicaragua, by SMITH & TAYLOR 1950

notatus: Hispaniola, including lIe à Cabrit, Ile de la Gonâve; Jamaica; Grand Cayman Island. Altitudinal distribution sea level to 3000 ft.; Type locality: Aquin, Departement du Sud, Haiti (but see comment).

Type locality: Martinique and Cuba.  
TypesSyntypes: MNHN-RA 1766 and MNHN-RA 1766A-B
Holotype: ZMUC (also as UZM) R.34462 [notatus]
Holotype: unlocated, fide Schartz & Henderson 1988 [fuscus]
Holotype: MBUCV 8.291 (Museo de Biología Universidad Central de Venezuela), adult male, collected 1963 by the Rivero-Blanco. [bodinii] 
DiagnosisDiagnosis: The combination of lacking moveable eyelids and having the dorsal surface of the head covered with granular scales distinguishes Gonatodes albogularis from all other Honduran lizards, except the remaining Gekkonoidea. Aristelliger, Hemidactylus, and Thecadactylus have distinctively expanded subdigital lamellae and have vertically elliptical pupils (versus all lamellae narrow and pupil circular in G. albogularis). Phyllodactylus have the terminal subdigital lamella expanded, divided, forming paired leaflike pad, and have vertically elliptical pupils (versus terminal lamella narrow, but divided, and pupils circular in G. albogularis). Sphaerodactylus have the claws displaced laterally by expanded terminal subdigital lamella and have a spinelike superciliary scale (versus claws located medially between pair of divided, nonexpanded lamella and spinelike superciliary scale usually absent in G. albogularis) (McCranie 2018: 87). 
CommentIllustrations: see Stejneger, 1917; Barbour and Ramsden, 1919; Cochran, 1941; Smith, 1946; Conant and Collins 1991; Petzold, 1965.

Subspecies: Gonatodes albogularis fuscus (HALLOWELL 1855) is considered as a valid species by several authors.

Distribution: Not listed for Belize by KÖHLER (2000). Schartz & Henderson 1988 provide two different type localities for G. a. notatus, namely the one given above, and “Port-au-Prince, Petionville, Jeremie, Diquini, Miragoane, Petit-Goave, Croix des Bouquets, Gonalves, 12.8 mi. N Cavaillon)”. Vanzolini and Williams 1962 commented upon the questionable status of the type-material of G. a. albogularis from Martinique, from where the species has not been re-recorded. The nominate subspecies is known from South America (Vanzolini and WiIIiams 1962). Records from Trinidad and Tobago are dubious (Murphy 1997) and may represent G. vittatus which is fairly ubiquitous there (Stevland Charles, pers. comm. 1 March 2018). For maps of species distributions see Vanzolini 1968: 89 (Map 1).

Type species: Gymnodactylus albogularis DUMÉRIL & BIBRON 1836 is the type species of the genus Gonatodes FITZINGER 1843: 91. Fitzinger (1843) defined the genus Gonatodes to accommodate three species, G. gravenhorstii, G. laevis, and G. albogularis but without designating a type species. With gravenhorstii and laevis being nomina nuda, albogularis becomes the type species (see Rösler et al. 2019 for a discussion).

Description: Gonatodes species are generally sexually dichromatic, with males being conspicuously ornamented with bright colors, whereas females are always cryptically colored (Rivero-Blanco 1979). The only exception seems to be Gonatodes lichenosus.

Behavior: mainly diurnal 
EtymologyThe name albogularis is derived from the Latin words albus (white) and gula (throat) and Latin suffix -arius (pertaining to). The name refers to the white throats of the syntypes alluded to by A. M. C. Duméril and Bibron (1836). 
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