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Gonocephalus megalepis (BLEEKER, 1860)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Bleeker's Forest Dragon 
SynonymLophyrus megalepis BLEEKER 1860: 327
Tiaris tuberculatus GÜNTHER 1872 (fide BOULENGER 1887)
Gonyocephalus tuberculatus — BOULENGER 1885: 291
Gonyocephalus megalepis — BOULENGER 1887: 152
Gonyocephalus megalepis — DE ROOIJ 1915: 107
Gonocephalus megalepis — WERMUTH 1967: 60
Gonocephalus megalepis — MANTHEY & GROSSMANN 1997: 188
Gonocephalus megalepis — TEYNIÉ et al. 2010
Gonocephalus megalepis — HARVEY et al. 2021 
DistributionIndonesia (Sumatra)

Type locality: Agam, Sumatra.

Type locality: “East Indian Archipelago” [Tiaris tuberculatus]  
TypesHolotype: BMNH 1946.8.27.21, male, (= holotype of Tiaris tuberculatus), Dr. Bleeker, apparently also the holotype of megalepis (fide W. Denzer, pers. comm., 18 June 2019)
CommentSynonymy mostly after WERMUTH 1967.

Distribution: see map in Harvey et la. 2021: 106 (Fig. 7). 
  • Bleeker, P. 1860. Reptilien van Agam. Natuurkundig Tijdschrift voor Nederlandsch Indie, Batavia, 20: 325-329 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, G.A. 1887. Note on some reptiles from Sumatra described by BLEEKER in 1860. Ann. Mag. Nat. Hist. (5) 20: 152 - get paper here
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Günther, A. 1872. On the reptiles and amphibians of Borneo. Proc. Zool. Soc. London 1872: 586-600. - get paper here
  • Harvey, Michael B.; Inna Rech, Awal Riyanto, Nia Kurniawan & Eric N. Smith 2021. A New Angle-Headed Dragon (Agamidae: Gonocephalus) from Montane Forests of the Southern Bukit Barisan Range of Sumatra Herpetologica Mar 2021 Vol. 77, No. 1: 94-109 - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Manthey U; Denzer W 1991. Die Echten Winkelkopfagamen der Gattung Gonocephalus Kaup (Sauria: Agamidae). 1. Die megalepis-Gruppe mit Gonocephalus lacunosus sp. n. aus Nord-Sumatra. Sauria 13 (1): 3-10 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
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