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Grayia smithii (LEACH, 1818)

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Higher TaxaColubridae, Grayiinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Smith's African Water Snake 
SynonymColuber smythii LEACH 1818: 409
Coluber laevis HALLOWELL 1844
Coronella triangularis — HALLOWELL 1854
Grayia silurophaga — GÜNTHER 1858: 51
Grayia triangularis BOCAGE 1866: 47
Graya [sic] silurophaga — ROCHEBRUNE 1884 (fide PAUWELS & MEIRTE 1996)
Grayia silurophaga — DOLLO 1887
Grayia triangularis — GÜNTHER 1888: 325
Grayia smythii — GÜNTHER 1895: 525
Grayia smythii — SCHMIDT 1923: 94
Grayia smythii — LOVERIDGE 1955
Grayia smythii — PITMAN 1974
Grayia smithi — GRUSCHWITZ et al. 1991 (fide ZR)
Grayia smithii — MEIRTE 1992: 113
Grayia smithii — TRAPE & ROUX-ESTÈVE 1995: 40
Grayia smythii — RÖDEL et al. 1999
Grayia smythi — RÖDEL & MAHSBERG 2000
Grayia smythii — SPAWLS et al. 2001
Grayia smithii — BRANCH & RÖDEL 2003
Grayia smithi — TRAPE & MANE 2004
Grayia smithi — TRAPE & BALDÉ 2014
Grayia smythii — WALLACH et al. 2014: 316
Grayia smithii — SPAWLS et al. 2018: 536
Grayia smythii — SANTOS et al. 2021 
DistributionRepublic of South Sudan (RSS), Uganda, Kenya, Tanzania, Rwanda, Burundi, Angola, Democratic Republic of the Congo (Zaire), Congo (Brazzaville), Equatorial Guinea, Cameroon, Nigeria, Benin, Togo, Ghana, Ivory Coast, Liberia, Guinea (Conakry), Guinea-Bissau, Senegal, Burkina Faso, Sierra Leone, Gambia, Central African Republic, probably S Mali (Joger & Lambert 1996), Niger

Type locality: Embomma = Boma, Zaire.  
Reproductionoviparous 
TypesType: BMNH 1946.1.1.67 (and possibly additional specimens). 
Diagnosis 
CommentNot in Gabon fide PAUWELS & VANDE WEGHE 2008.

Type species: Coluber smythii LEACH 1818: 409 is the type species of the genus Grayia GÜNTHER 1858: 50. Grayia is also the type genus of the subfamily Grayiinae Meirte 1992. 
EtymologyNamed after Norwegian botanist Christen Smith (1785-1816); hence the name “smythii” is inappropriate (but see Laurent 1956 and Meirte 1992). 
References
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