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Leiocephalus barahonensis SCHMIDT, 1921

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Higher TaxaLeiocephalidae, Iguania, Sauria, Squamata (lizards)
SubspeciesLeiocephalus barahonensis barahonensis SCHMIDT 1921
Leiocephalus barahonensis altavelensis NOBLE & HASSLER 1933
Leiocephalus barahonensis aureus COCHRAN 1934
Leiocephalus barahonensis beatanus NOBLE 1923
Leiocephalus barahonensis oxygaster SCHWARTZ 1967 
Common NamesE: Orange-bellied curlytail, Barahona Curlytail Lizard 
SynonymLeiocephalus barahonensis SCHMIDT 1921:15
Leiocephalus personatus barahonensis — MERTENS 1938
Leiocephalus personatus barahonensis — MERTENS 1939
Leiocephalus barahonensis — SCHWARTZ 1967
Leiocephalus barahonensis — SCHWARTZ & HENDERSON 1991: 421
Leiocephalus barahonensis — PREGILL 1992: 38
Leiocephalus barahonensis — PYRON 2017

Leiocephalus barahonensis altavelensis NOBLE & HASSLER 1933
Leiocephalus altavelensis NOBLE & HASSLER 1933: 14
Leiocephalus vinculum altavelensis — SCHWARTZ 1967
Leiocephalus vinculum altavelensis — SCHWARTZ & HENDERSON 1988: 150
Leiocephalus barahonensis altavelensis — PREGILL 1992
Leiocephalus altavelensis — HEDGES et al. 2019

Leiocephalus barahonensis aureus COCHRAN 1934
Leiocephalus personatus aureus COCHRAN 1934: 175
Leiocephalus barahonensis aureus — SCHWARTZ & HENDERSON 1988: 135
Leiocephalus barahonensis aureus — HEDGES et al. 2019

Leiocephalus barahonensis beatanus NOBLE 1923
Leiocephalus beatanus NOBLE 1923: 555
Leiocephalus personatus beatanus — COCHRAN 1934: 177
Leiocephalus barahonensis beatanus — SCHWARTZ & HENDERSON 1988: 135
Leiocephalus personatus beatanus — TIEDEMANN et al. 1994: 51
Leiocephalus personatus beatanus — HEDGES et al. 2019

Leiocephalus barahonensis oxygaster SCHWARTZ 1967
Leiocephalus barahonensis oxygaster SCHWARTZ 1967
Leiocephalus barahonensis oxygaster — SCHWARTZ & HENDERSON 1988: 135
Leiocephalus barahonensis oxygaster — HEDGES et al. 2019 
DistributionSC Hispaniola (SE Haiti, islands off east coast of Peninsula de Barahona)

barahonensis: Repûblica Dominicana, north slope of Sierra de Baoruco to northeast coast of Peninsula de Barahona. Elevation: sea level to 1000 ft. Type locality: Barahona, Barahona Province, República Dominicana.

altavelensis: Isla Alto Velo. Type locality: Isla Alto Velo, Republica Dominicana.

aureus: extreme southeastern Haiti, east to Dominico-Haitian border; intergrades with L. b. oxygaster SE Pedernales, República Dominicana. Elevation: sea level to 1900 ft.

beatanus: Isla Beata. Type locality: Isla Beata, Republica Dominicana.

oxygaster: Repéblica Dominicana, Peninsula de Barahona; presumed subspecies on Cayo Pisaje. Elevation: near sea level. Type locality: “Type locality: 13.1 mi. (21.1 km) SW Enriquillo, Pedernales Province, Republica Dominicana.  
TypesHolotype: AMNH 2736.
Holotype: AMNH 51055 [altavelensis]
Holotype: AMNH 24330, Paratype: NMW 16621 [beatanus]
Holotype: USNM 75909 [aureus]
Holotype: MCZ 81098 [oxygaster] 
DiagnosisDESCRIPTION: Size moderate (SVL in males to 80 mm, in females to 64 mm); sexual dichromatism not pronounced dorsally; throat pattern somewhat variable between sexes; males with 2 pairs of enlarged postanals; no lateral fold; dorsals imbricate, strongly denticulate or tricuspid, keeled; ventrals imbricate, smooth, denticulate; median dorsal crest scales slightly enlarged, not attenuate, slightly imbricate, slightly lower than median dorsal caudal scales, 42-57 in occiput-vent distance; one-half midbody scales 16-24; supraoculars usually 6/6; loreals 2-8; temporals 3-7; supraorbital semicircles always incomplete; parietals always in contact; median head scales 2~; preauricular scale extremely enlarged. Throat in both sexes with gray to black reticulum, black transverse bars, or dark smudges; venter bright orange or yellow-orange, patternless in both sexes; facial mask absent; neck and shoulder patches absent; dorsum with a broad zone of golden yellow, slightly reddish, brown, ashy gray, brownish orange, vivid orange-red, with or without an included pair of paramedian (plus a median in 1 subspecies) dorsal stripes; light dorsolateral stripes distinct to indistinct; throat dark (gray, dirty orange, gray with bronzy sheen) to pale (cream, white) (Schwartz & Henderson 1991: 422). 
CommentFor illustrations see Noble, 1923 (as L. beatanus); Cochran, 1941 (as subspecies of L. personatus); Schwartz, 1967. 
  • Cochran, D. M. 1934. Herpetological collections made in Hispaniola by the Utowana Expedition. Occ. Pap. Boston Soc. Nat. Hist. 8: 163-188 - get paper here
  • Cochran, D.M. 1941. The herpetology of Hispaniola. Bull. US Natl. Mus. 177: vii + 398 pp. - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Hedges SB, Powell R, Henderson RW, Hanson S, and Murphy JC 2019. Definition of the Caribbean Islands biogeographic region, with checklist and recommendations for standardized common names of amphibians and reptiles. Caribbean Herpetology 67: 1–53
  • Mertens, R. 1938. Amphibien und Reptilien aus Santo Domingo, gesammelt von Prof. Dr. H. Böker. Senckenbergiana 20 (5): 332-342
  • Micco, Shonette M.;Lahey, Garrett J.;Sosa, Roland A.;Powell, Robert;Censky, Ellen J.;Parmerlee, John S. Jr 1998. Natural History of Leiocephalus barahonensis(Tropiduridae) on the Peninsula de Barahona, Hispaniona: an examination of two populations. Herpetological Natural History 5 (2): 147-156 [1997]
  • Noble, G. K. and W. G. Hassler 1933. Two new species of frogs, five new species and a new race of lizards from the Dominican Republic. American Museum Novitates 652: 1-17. - get paper here
  • Noble, G.K. 1923. Trailing the rhinoceros iguana. Nat. Hist. 23 (6):541-558.
  • Pregill,G.K. 1992. Systematics of the West Indian Lizard Genus Leiocephalus (Squamata: Iguania: Tropiduridae). Misc. Publ. Univ. Kans. Mus. Nat. Hist. 84: 1-69 - get paper here
  • Pyron, R. Alexander 2017. Novel Approaches for Phylogenetic Inference from Morphological Data and Total-Evidence Dating in Squamate Reptiles (Lizards, Snakes, and Amphisbaenians) Syst. Biol. 66 (1): 38–56 - get paper here
  • Schmidt, K. P. 1921. Notes on the herpetology of Santo Domingo. Bull. Amer. Mus. Nat. Hist. 44: 7-20. - get paper here
  • Schwartz, A. 1967. The Leiocephalus (Lacertilia: Iguanidae) of Hispaniola. II. The Leiocephalus personatus complex. Tulane Studies in Zoology 14: 1-53 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
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