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Liolaemus koslowskyi ETHERIDGE, 1993

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Higher TaxaLiolaemidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymLiolaemus koslowskyi ETHERIDGE 1993: 159
Liolaemus (Eulaemus) koslowskyi — SCHULTE et al. 2000
Liolaemus koslowskyi — CAMARGO et al. 2012
Liolaemus koslowskyi — AVILA et al. 2021 
DistributionArgentina (Catamarca, La Rioja)

Type locality: 9.2 km East de la plaza central de Pituil,
Ruta Pcial 11, Dpto Famatina.  
ReproductionOviparous. 
TypesHolotype: FML 2659 
DiagnosisDiagnosis. Liolaemus koslowskyi differs from all other members ofthe L. darwinii complex, except L. darwinii, L. abaucan and, perhaps L. uspallatensis in having conspicuously enlarged, dark brown or black prescapular and postscapular spots, from L. darwinii in lacking intensely dark pigment within the antehumeral fold, and from L. abaucan and L. uspallatensis in having larger, more strongly keeled dorsal scales and a different dorsal color pattern. (Etheridge 1993: 160).

Variation. Maximum size, and variation in tail/total length, scale counts and precloacal pore numbers provided in Tables 1-4. Males 41 to 48 mm snout-vent length. ( (Etheridge 1993: 160).

Coloration: Males with standard pattern, modified only by enlargement and darkening of prescapular and postscapular spots; smaller males unknown. Adult males characteristically with pattern as described for holotype: middorsal and dorsolateral light stripes prominent on neck and anterior body, fading to obscurity on posterior body and tail; prescapular and postscapular spots enlarged, very dark and conspicuous; paravertebral spots enlarged and diffuse, oftenjoined medially, when alternate forming a dark zig-zag with diffuse edges between dorsolateral stripes. Sides ofbody usually speckled, but one or more dark, lateral spots posterior to postscapular may be present. Wide, conspicuous, dorsolateral light stripes occasionally present from head to tail. Throat always spotted, but marks may be few and indistinct, or numerous and discrete, at times forming a reticulum; ventral body immaculate or with scattered, indistinct greyish spots; apparently no ontogenetic change in throat and ventral body pattern.
Females with standard pattern retained or reduced and faded, usually at least some indication of dorsolateral stripes and paravertebral spots remaining the stripes often prominent. Some females with postscapular spots somewhat enlarged and darkened. Throat and ventral body with same range ofvariation as males, but tendency for fewer, less distinct markings. Gravid females with orange or reddish-orange pigment on the sides of the head and neck. Populations on sandy flats such as Bajo de Santa Elena (PI. 1.4) and the borders of Salar de Pipanaco, with pale tan ground color; individuals inhabiting rocky areas darker (PI. 1.5). (Etheridge 1993: 160). 
CommentMember of the darwinii group of Liolaemus. 
EtymologyNamed to honor Julio Koslowsky (1886-1923), in recognition of his contributions to the herpetology of Argentina, and especially the provinces ofLa Rioja and Catamarca (Koslowsky, 1895, 1898), where this species occurs. 
References
  • Abdala, C.S. 2005. Una nueva especie del género Liolaemus perteneciente al complejo darwinii (Iguania:Liolaemidae) de la provincia de Catamarca, Argentina. Revista Española de Herpetologia 19:5-17 - get paper here
  • Abdala, C.S. 2007. Phylogeny of the boulengeri group (Iguania: Liolaemidae, Liolaemus) based on morphological and molecular characters. Zootaxa 1538: 1-84 - get paper here
  • Abdala, Cristian Simón; Quinteros, Andres Sebastian 2014. Los últimos 30 años de estudios de la familia de lagartijas más diversa de Argentina. Actualización taxonómica y sistemática de Liolaemidae. Cuad. herpetol. 28 (2): 55-82 - get paper here
  • ABDALA, CRISTIAN SIMÓN; ANDRÉS SEBASTIÁN QUINTEROS,, FEDERICO ARIAS, SABRINA PORTELLI & ANTONIO PALAVECINO 2011. A new species of the Liolaemus darwinii group (Iguania: Liolaemidae) from Salta Province, Argentina. Zootaxa 2968: 26–38 - get paper here
  • Avila, L. J., M. Morando, and D. R. Perez. 1999. Geographic distribution. Phymaturus patagonica. Herpetological Review 30 (3): 174 - get paper here
  • AVILA, LUCIANO JAVIER; LORENA ELIZABETH MARTINEZ & MARIANA MORANDO 2013. Checklist of lizards and amphisbaenians of Argentina: an update. Zootaxa 3616 (3): 201–238 - get paper here
  • Avila, Luciano Javier; Martín Miguel Femenías, Cristian Hernán Fulvio Pérez, Mariana Morando 2021. Nuevas especies de Liolaemus para la herpetofauna de la provincia de La Rioja, Argentina. Cuadernos de Herpetología 35 (S1): 223-228
  • Belver, Luciana C.; Kozykariski, Mónica L.; Avila, Luciano J. 2010. Diferencias sexuales y etarias en la actividad diaria y estacional de una población de Liolaemus koslowskyi (Liolaemini). Cuadernos de Herpetología 24 (2) - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Camargo, Arley; Luciano J. Avila, Mariana Morando, and Jack W. Sites, Jr 2012. Accuracy and Precision of Species Trees: Effects of Locus, Individual, and Base Pair Sampling on Inference of Species Trees in Lizards of the Liolaemus darwinii Group (Squamata, Liolaemidae). Systematic Biology 61: 272-288 - get paper here
  • Etheridge, R. 1993. Lizards of the Liolaemus darwinii complex (Squamata: Iguania: Tropiduridae) in Northern Argentina. MUSEO REGIONALE DI SCIENZE NATURALI BOLLETTINO (TURIN). 11 (1): 137-199
  • Etheridge, R. 2000. A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs 14: 293-352 - get paper here
  • Morando, M.; Avila, L.J.; Baker, J. & Sites, J.W. 2004. PHYLOGENY AND PHYLOGEOGRAPHY OF THE LIOLAEMUS DARWINII COMPLEX (SQUAMATA: LIOLAEMIDAE): EVIDENCE FOR INTROGRESSION AND INCOMPLETE LINEAGE SORTING. Evolution 58 (4): 842–861 - get paper here
  • SCHULTE II, JAMES A.; J. ROBERT MACEY, ROBERT E. ESPINOZA AND ALLAN LARSON 2000. Phylogenetic relationships in the iguanid lizard genus Liolaemus: multiple origins of viviparous reproduction and evidence for recurring Andean vicariance and dispersal. Biological Journal of the Linnean Society 69: 75–102 - get paper here
  • Tulli, M.J.; F.B. Cruz, A. Herrel, B. Vanhooydonck, V. Abdala 2009. The interplay between claw morphology and microhabitat use in neotropical iguanian lizards. Zoology 112 (5): 379-392 - get paper here
 
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