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Nactus allenallisoni ZUG, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Madang Slender-toed Gecko 
SynonymNactus allenallisoni ZUG 2020: 42 
DistributionN Papua New Guinea (from Aitape eastward to Guisko and Finschhafen of coastal Huon Peninsula and also on Manam and Karkar Islands)

Type locality: Papua New Guinea, Madang Province, Siar Plantation [−5.1917° 145.7774°], Alexishafen  
Reproduction 
TypesHolotype. AMS R31261, adult male, collected by S. Donnellan on 19 March 1987 (Figure 16).
Paratypes. AMNH 104874, adult male from Wanuma, Adelbert Mountains [−4.9195° 145.3182°], Madang Province, collected by R. G. Zweifel on 4 August 1969; AMNH 104873, adult female, vicinity of Sempi [−5.1486° 145.7715°], ca. 5 miles [~8 km] N of Alexishafen, Madang Province collected by R. G. Zweifel on 13 July 1969; AMS R31260, R31262, R31268, R31281, adult males with same collecting data as holotype; AMS R31272, R31284, R31291, R31297, adult females with same collecting data as holotype. 
DiagnosisDefinition. A bisexual taxon of geckos (Gek- koninae) with moderate to large adults (♀♀ 47–64 mm, ♂♂ 46–63 mm SVL) with keeled subcaudals, variable-sized postmen- tal scales (0–22, median 14), no or slight postmental-infralabial (PmLab) contact, low to moderate number of dorsal tubercle rows (12–22) and moderate number of tubercles (22–36, usu- ally ≥28) in parasagittal row (TubRow), tubercles on dorsal surface of thigh and crus, and few precloacal pores (2, 0–9) in males (most females without pores, 0–3). Diagnostic summary in Table 8.

Coloration in life unknown. In alcohol, background rufous tan dorsally and lighter ventrally. Top of head with scattered, smudge-like medium-brown marks. Dorsally dark brown, para- sagittal smudge-like marks on middle of neck, 5 pairs of parasag- ittal dark-brown blotches on trunk and 1 on base of tail. Limbs with scattered diffuse dark marks.

Variation. Females and males display no striking metric differences (adult ♀♀ 56.6, 47.0–64.3 mm; adult ♂♂ 53.2, 45.9–62.5 mm SVL) and also share proportions (total adult sample) TrunkL/SVL 39.8%, 32%–46%; HeadL/SVL 27.4%, 25%–40%; HeadW/SVL 18.1%, 15%–22%; HeadW/HeadL 65.9%, 48%–75%; EyeD/SVL 6.7%, 5%–9%; EyeD/HeadL 24%, 19%–33%; NarEye/HeadL 32.2%, 21%–43%; Interorb/ HeadL 27.1%, 21%–32%; SnW/HeadL 14.5%, 9%–18%.
Typical head scale pattern of small, granular scales except for those bordering nares and mouth. Snout with large rostral scale bearing middorsal cleft posteriorly; large supralabials (3, 3–4) and infralabials (3, 3–4) in front of anterior edge of orbit, first supra- and infralabial largest of respective series. Chin with large trapezoidal mental and usually modest to large postmentals (Postm = 14, 0–22, single specimen lacking postmental) and usually no genial scale between postmental and first surpalabial (0, 0–3 PmLab); dorsally, trunk with 17, 12–22 (DorsTub) enlarged tubercles transversely on dorsum and with 29, 22–36 tubercles (TubRow) longitudinally along trunk; 10, 8–10 tubercle rows transversely between hindlimbs (TubHip). Scattering of large tubercules on dorsal half of fore- limb and more on hindlimb (TubHindl = 2). Tail with small uniform scales dorsally and laterally, ventrally with uniform, small keeled scales (Subcaud = 0, with exception of one indi- vidual with smooth subcaudals). Usually, 1 cloacal spur (1, 0–2 CloacS) on each side. Ventral scales from chin to vent small, granular, somewhat larger on chest and abdomen. Precloacal pores, males 0–9, females 0–3. Forefoot with narrow digits, their lengths usually 3>4≈2>5≈1; 17, 15–20 lamellae beneath fourth digit (4FingLm), 5, 4–7 scale rows on palm; hindfoot with narrow digits, their lengths 4≈3>5>2>1; 22, 19–25 lamel- lae beneath fourth toe (4ToeLm).
Coloration in life not reported. In alcohol, dorsal back- ground medium brown from snout onto tail, ill-defined inter- orbital dark-brown mark extending narrowly onto snout, dark-brown reverse triangular mark on parietal-crown, and series of paired parasagittal dark-brown marks from nape onto base of tail; brown of dorsum grading into lighter brown of ven- ter, uniform from chin onto base of tail. 
CommentSympatry: N. septentrionalis in Aitape, Nuku, Wewak, and Manam Island; N. nanus at Siar Plantation

Distribution: see map in Zug 2020: 47 (Fig. 18). 
EtymologyThis species is named to recognize Allen Allison’s contributions to the biology of the New Guinea herpe- tofauna and his continuing role in the conservation of the Papua New Guinea biota by encouraging and supporting the active involvement of PNG villagers and tribes. The specific name is a noun in apposition. 
References
  • Zug, George R. 2020. Diversity in Pacific Slender-Toed Geckos, Nactus pelagicus Complex (Reptilia: Squamata), of New Guinea and Adjacent Islands. Smithsonian Contributions to Zoology, (651): x + 92 pages - get paper here
 
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