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Nactus amplus ZUG, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Louisiade Giant Slender-toed Gecko 
SynonymNactus amplus ZUG 2020: 58 
DistributionPapua New Guinea (Milne Bay Province: Louisiade Archipelago: Sudest Island, Rossel (Yela) Island)

Type locality: Papua New Guinea, Milne Bay Province, Rossel Island, track up Mount Rossel, Point 30, small stream below Camp 2, −11.3544° 154.2232°E.  
Reproduction 
TypesHolotype. BPBM 19864, adult female, collected by Fred Kraus, F. Malesa, and local collectors on 9 May 2004 (Figure 26).
Paratypes. Adult females from Rossel Island, Louisiade Archipelago, Milne Bay Province, Papua New Guinea, AMNH 76752, Mount Rossel, south slope, 700 m, collected by R. F. Peterson on 12–20 October 1956; BPBM 19867, Rossel Island, Camp 3 along Rupu River, −11.3354° 154.2247°, col- lected by Fred Kraus, F. Malesa, and local collectors on 12 May 2004, and immature female, BPBM 19869, Wupu River crossing, −11.3380° 154.2238°, 278 m, collected by Fred Kraus, F. Malesa, and local collectors on 12 May 2004. Adult females, BPBM 19847, 19848 from Sudest Island, Camp 1, −11.4918° 153.4162°, 127 m elevation, along Gesirava River, collected by Fred Kraus on 16 April 2004. 
DiagnosisDefinition. A bisexual taxon of geckos (Gekko- ninae) with large adults (♀♀ 75–82 mm, ♂ 68 mm SVL) with smooth subcaudals, variable-sized postmental scales (6–24, median 12), variable postmental-infralabial (PmLab = 0–2, median 0) contact, moderate number of dorsal tubercle rows (14–20) and moderate number of tubercles (27–32, usually ≥29) in parasagittal row (TubRow), tubercles usually present on dorsal surface of thigh and crus, and precloacal pores variable (median 0, 0–14 in females; 14 in male). Diagnostic summary in Table 8.

Variation. Adult females range between 75.5 and 82.1 mm SVL, adult male 67.9 mm SVL; the smallest individual in the sample is an immature female of 66.7 mm SVL. For adults, TrunkL/SVL 41.3%, 39%–44%; HeadL/SVL 28.4%, 27%– 30%; HeadW/SVL 18.8%, 17%–20%; HeadW/HeadL 64.8%, 62%–69%; EyeD/SVL 7.1%, 7%–9%; EyeD/HeadL 24.9%, 23%–28%; NarEye/HeadL 32.8%, 31%–34%; Interorb/HeadL 25.6%, 24%–27%; SnW/HeadL 13.1%, 12%–13%.
Typical head scale pattern of small, granular scales except for those bordering nares and mouth. Snout with large rostral scale bearing middorsal cleft posteriorly; supranasals moderate sized, usually in contact on midline or separated by small granu- lar scale (internasal); large supralabials (4, 3–4) and infralabi- als (3, 3–4) in front of anterior edge of orbit, first supra- and infralabial largest of respective series. Chin with large trapezoi- dal mental and small to moderately large postmentals (Postm = 12, 6–24) and usually no genial scale between postmental and first supralabial (0, 0–2 PmLab); dorsally, trunk with 17, 14–20 (DorsTub) enlarged tubercles transversely on dorsum and with 29, 27–32 tubercles (TubRow) longitudinally along trunk; 10, 8–10 tubercle rows transversely between hindlimbs (TubHip). Enlarged tubercles on crus and femur (TubHindl = 2). Tail with small uniform scales dorsally and laterally, ventrally typically with uniform, small keeled smooth (Subcaud = 1). Usually, pair of cloacal spurs (2, 2–3 CloacS) on each side. Ventral scales from chin to vent small, granular, somewhat larger on chest and abdomen. Precloacal pores, male 14, females usually none, 0–11. Forefoot with narrow digits, their lengths usually 3≈4>2≈5>1; 19, 18–21 lamellae beneath fourth digit (4FingLm), 5, 5–7 scale rows on palm; hindfoot with narrow digits, their lengths 3≈4>2≈5>1; 23, 23–25 lamellae beneath fourth toe (4ToeLm).
Coloration in life not reported. In alcohol, color and pattern match the general description of the holotype. 
CommentDistribution: see map in Zug 2020: 53 (Fig. 22). 
EtymologyThe Latin amplus refers to “large.” This population contains the largest known adults in the genus Nactus. 
References
  • Zug, George R. 2020. Diversity in Pacific Slender-Toed Geckos, Nactus pelagicus Complex (Reptilia: Squamata), of New Guinea and Adjacent Islands. Smithsonian Contributions to Zoology, (651): x + 92 pages - get paper here
 
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