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Nactus fredkrausi ZUG, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Kraus’s Giant Slender-toed Gecko 
SynonymNactus fredkrausi ZUG 2020: 54 
DistributionPapua New Guinea (montane forest of central Misima Island)

Type locality: Papua New Guinea, Milne Bay Province, Misima Island, Oya Tau, “Camp 2,” 860 m asl, 10°39.355′S 152°37.659′E.  
Reproduction 
TypesHolotype. BPBM 16749, adult male, collected by Fred Kraus on 18 January 2003 (Figure 23).
Paratypes. BPBM 16742, 16746, 16751, 16753, adult females, and BPBM 16745, 16750, 16752, adult males; all from Misima Island and collected by Fred Kraus. BPBM 16742, 16745–746 from Bwaga Bwaga ridge camp, −10°42.437′ 152°40.971′, 350–400 m, collected on 12 January 2003; BPBM 16750–753 with the same collecting data as the holotype. 
DiagnosisDefinition. A bisexual taxon of geckos (Gekko- ninae) with large adults (♀♀ 57–77 mm, ♂♂ 52–74 mm SVL) with keeled subcaudals, variable-sized postmental scales (8–22, median 14), no or slight postmental-infralabial contact (PmLab = 0–1, median 0), moderate number of dorsal tubercle rows (16–19) and moderate number of tubercles (26–33, usually ≥29) in para- sagittal row (TubRow), tubercles present on dorsal surface of thigh and crus, and precloacal pores present in males (4–7), usu- ally present in females (0–7). Diagnostic summary in Table 8.

Variation. Females and males display slight met- ric differences (adult ♀♀ 67.6, 57.1–76.5 mm; adult ♂♂ 66.1, 52.0–74.2 mm SVL) and share similar proportions (total adult sample): TrunkL/SVL 39.0%, 33%–42%; HeadL/SVL 28.7%, 28%–30%; HeadW/SVL 17.0%, 15%–18%; HeadW/HeadL 59.3%, 53%–66%; EyeD/SVL 7.1%, 7%–8%; EyeD/HeadL 24.8%, 23%–28%; NarEye/HeadL 33.9%, 32%–34%; Inter- orb/HeadL 25.0%, 22%–28%; SnW/HeadL 13.8%, 13%–15%.
Typical head scale pattern of small, granular scales except for those bordering nares and mouth. Snout with large rostral scale bearing middorsal cleft posteriorly; supranasals moderate sized, usually in contact on midline or separated by small granu- lar scale (internasal); large supralabials (3, 3–4) and infralabi- als (3, 3–4) in front of anterior edge of orbit, first supra- and infralabial largest of respective series. Chin with large trapezoi- dal mental and usually moderate-sized postmentals (Postm = 14, 8–22) and usually no genial scale between postmental and first supralabial (0, 0–1 PmLab); dorsally, trunk with 18, 16–19 (DorsTub) enlarged tubercles transversely on dorsum and with 29, 26–33 tubercles (TubRow) longitudinally along trunk; 10, 8–11 tubercle rows transversely between hindlimbs (TubHip). Uniform covering of small tuberculate-like scale on dorsal half of forelimb and enlarged tubercles on femur (TubHindl = 1). Tail with small uniform scales dorsally and laterally, ventrally typi- cally with uniform, small keeled scales (Subcaud = 0). Usually, paired cloacal spurs (2, 0–4 CloacS) on each side. Ventral scales from chin to vent small, granular, somewhat larger on chest and abdomen. Precloacal pores, males 4–7, females 0–6, most with pores. Forefoot with narrow digits, their lengths usually 3≈4>2>5>1; 19, 17–21 lamellae beneath fourth digit (4FingLm), 6, 5–7 scale rows on palm; hindfoot with narrow digits, their lengths 4≈3>2≈5>1; 24, 21–26 lamellae beneath fourth toe (4ToeLm).

Coloration in life not reported. In alcohol, pattern and col- oration largely match holotype. 
CommentDistribution: see map in Zug 2020: 53 (Fig. 22). 
EtymologyThis species is named to recognize Fred Kraus’s past and continuing contributions to the systemat- ics and biology of the New Guinea herpetofauna. I also wish to acknowledge our friendship and his ongoing tolerance for answering my questions on New Guinea and the exotic herpe- tofauna of Hawaii. The specific name is a noun in apposition. 
References
  • Zug, George R. 2020. Diversity in Pacific Slender-Toed Geckos, Nactus pelagicus Complex (Reptilia: Squamata), of New Guinea and Adjacent Islands. Smithsonian Contributions to Zoology, (651): x + 92 pages - get paper here
 
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