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Oxybelis inkaterra JADIN, JOWERS, ORLOFSKE, DUELLMAN, BLAIR & MURPHY, 2021

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Inkaterra Vine Snake
S: Inkaterra Serpiente de vid 
SynonymOxybelis inkaterra JADIN, JOWERS, ORLOFSKE, DUELLMAN, BLAIR & MURPHY 2021
Oxybelis aeneus – KEISER, 1974:7;
Oxybelis aeneus – DUELLMAN 2005: 363, pl 205
Oxybelis aeneus – JADIN et al. 2020: Fig 1. 
DistributionPeru (Madre de Dios)

Type locality: Peru, Madre de Dios: Cusco Amazónico (now Reserva Amazónica), Río Madre de Dios, c. 15 km E Puerto Maldonado, 200 m, 12°34'59"S, 69°4'59"W  
Reproduction 
TypesHolotype. KU 220196 (Figs 5, 6), collected by William E. Duell- man (WED 59561), 21 December 1991.
Paratypes. KU 214887, Peru, Madre de Dios: Cusco Amazónico, Río Madre de Dios, c. 15 km E Puerto Mal- donado, 200 m, 12°34'59.88"S, 69°4'59.879"W; collect- ed by Erik R. Wild (Field number WED 59004), 23 De- cember 1989; ZMH R01702, Peru, Huánuco: Pachitea, Panguana Biological Fieldstation, Rio Yuyapichis [= Rio Llullapichis], 260 m, approx. 9°41'S, 74°57'W, collected by János Regös July 1980. 
DiagnosisDiagnosis. A vine snake with (1) three upper labials (4–5–6) bordering the orbit; (2) numerous bold black bars and spots present on the body; (3) ventral surface mottled with dense black spots; (4) preocular shorter than eye diameter; (5) second pair of chin shields separated by smaller scales posteriorly; (6) nine upper labials, three located behind the orbit; (7) snout from above relatively broad, tapered, and flat rostrum; (8) supraocular and prefrontal are about the same length; (9) last upper labial about same length as primary temporal; (10) much of the lower surface of the head infused with black pigment; (11) second upper labial not in contact with preocular.

Comparison. Oxybelis inkaterra can be distinguished from the seven other members of the Oxybelis aeneus complex by the presence of, upper labials three and four are in contact the preocular; a head with an irregular, darkly pigmented ventral surface with pale spots; and eyespot markings on the posterior ventral surface of the body and tail; the snout of O. inkaterra is also relatively short and broad compared to other species in the Oxybelis aeneus complex (Fig. 7); the lack the brown-gray dorsal coloration seen in the other members of the complex. The dorsum is instead a dirty cream with black flecking (Figs 6, 8). 
CommentDetails and references not provided by authors upon request. 
EtymologyThe specific epithet honors the ecotourism company Inkaterra (https://www.inkaterra.com/) and its non-profit NGO counterpart Inkaterra Asociación. These two institutions started in 1975 and 1978, respectively, were founded by José E. Koechlin von Stein to promote education and conservation of Peruvian culture and ecosystems. Inkaterra and Mr. Koechlin have been recognized numerous times with awards and accolades for providing sustainable ecotourism and research opportunities for scientists. The type locality, Cusco Amazónico (now Reserva Amazónica), is owned and operated by Inkaterra and is the site of one of the most thoroughly studied areas in the Neotropics, particularly for amphibian and reptile natural history (e.g. Duellman, 2005; Orlofske et al. 2012). 
References
  • Jadin RC, Jowers MJ, Orlofske SA, Duellman WE, Blair C, Murphy JC 2021. A new vine snake (Reptilia, Colubridae, Oxybelis) from Peru and redescription of O. acuminatus. Evolutionary Systematics 5(1): 1-12 - get paper here
 
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