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Pachydactylus carinatus BAUER, LAMB & BRANCH, 2006

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Richtersveld Gecko 
SynonymPachydactylus carinatus BAUER, LAMB & BRANCH 2006: 673
Pachydactylus serval onscepensis — MCLACHLAN & SPENCE 1966: 155
Pachydactyluscf. serval — BAUER & BRANCH 2003: 133
Pachydactylus carinatus — MASHININI & MAHLANGU 2013
Pachydactylus carinatus — BATES et al. 2014: 129 
DistributionRepublic of South Africa (Northern Cape Province), Namibia (Lüderitz, Karasburg Districts). Elevation 40-720 m.

This species is distributed throughout the Richtersveld National Park and in adjacent parts of southern Namibia (Figs. 21, 30–31 in Bauer et al. 2006). It occurs in areas along the Orange River both to the east and west of the park boundaries — Annisfontein in the west and several localities in the east, at least as far upstream as Goodhouse, where it is replaced by P. montanus. It extends southwards to about the level of Kuboes and northwards as far as Namuskluft in the west and Ai-Ais in the east.

Type locality: South Africa, Northern Cape Province, Richtersveld National Park, 13.3 km E of Oenna Mine (28°05′11′′S, 17°07′45′′E).  
TypesHolotype: CAS 201908 (Fig. 101): Adult female; coll. A.M. Bauer, 4 July 1996. paratypes: CAS, PEM, TM 
DiagnosisDIAGNOSIS.—A moderately sized species, to 45.7 mm SVL (CAS 201908). Pachydactylus carinatus may be distinguished from all other members of the P. serval/weberi group by the combination of the following characters: snout weakly inflated laterally; rostral enters nostril; supranasals in variable contact; scales on dorsum of head granular, flattened to very weakly domed, those on snout much larger than those of interorbital and parietal regions; very few small (2–3 times size of granules), round, conical tubercles on interorbital and parietal regions; dorsal scalation strongly heterogeneous, with small, oval, keeled tubercles arranged in 16 regular rows; no tubercles on thighs; toes moderately long, toe pads relatively narrow; typically five undivided lamellae beneath digit IVof pes; tail to at least 114% of SVL, strongly annulate, bearing whorls of moderately to very strongly keeled, pointed tubercles, well separated from each other; adult pattern of moderately small, irregular brown spots or larger markings more-or-less evenly distributed across light brown to grayish-brown dorsum, with some trace of a pale, dark-edged band across occiput and nape in some specimens; tubercular keels whitish to pale yellow, contrasting with darker spots on dorsum (Figs. 101–103 in Bauer et al. 2006); juveniles with dark brown to blackish body, with an wide, dark-edged ashy nape band and an thick ashy band covering lumbar and sacral regions as well as hindlimbs (Figs. 104–105 in Bauer et al. 2006), tail a bright orange (Fig. 105 in Bauer et al. 2006; see also Bauer and Branch 2003:133). Although similar to the juvenile pattern of P. serval, in the latter species the tail is dark rather than bright, the pale sacral area extends further anteriorly than in P. carinatusand the neck band is also broader.
CommentHABITAT: P. carinatus is rupicolous and occupies retreats under overhanging rock flakes and narrow cracks and crevices within and between rocks in bouldery areas. 
EtymologyETYMOLOGY.—The specific epithet carinatusis Latin for keeled and is in reference to the prominent keeled tubercles typical of this species. 
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bauer, Aaron M. Lamb, Trip. Branch, William R. 2006. A revision of the Pachydactylus serval and P. weberi groups (Reptilia: Gekkota: Gekkonidae) of Southern Africa, with the description of eight new species. Proc. Cal. Acad. Sci. 57 (12-24):595-709. - get paper here
  • CONRADIE, WERNER; WILLIAM R. BRANCH, & GILLIAN WATSON 2019. Type specimens in the Port Elizabeth Museum, South Africa, including the historically important Albany Museum collection. Part 2: Reptiles (Squamata). Zootaxa 4576 (1): 001–045 - get paper here
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Mashinini, P. L. and Mahlangu, L. M. 2013. An annotated catalogue of the types of gekkonid lizards (Reptilia: Squamata: Gekkonidae) in the Herpetology collection of the Ditsong National Museum of Natural History, South Africa. Annals of the Ditsong National Museum of Natural History 3: 165-181
  • Mclachlan, G.R.; Spence, J.M. 1966. The Genus Pachydactylus (Part 1). Ann. Cape Prov. Mus. 5: 149-156
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