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Pachydactylus rugosus SMITH, 1849

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Common Rough Gecko, Wrinkled Thick-toed Gecko, Rough Thick-toed Gecko, Rough-scaled gecko 
SynonymPachydactylus rugosus SMITH 1849
Pachydactylus rugosus — BOULENGER 1885: 204
Pachydactylus rugosus frater HEWITT 1935: 316 (fide MERTENS 1955)
Pachydactylus rugosus — FITZSIMONS & BRAIN 1958
Pachydactylus rugosus — WERMUTH 1965: 122
Pachydactylus rugosus rugosus — AUERBACH 1987: 85
Pachydactylus rugosus — KLUGE 1993
Pachydactylus rugosus — RÖSLER 2000: 99
Pachydactylus rugosus — BAUER 2013 (plates)
Pachydactylus rugosus — CONRADIE et al. 2019 
DistributionRepublic of South Africa (S Little Namaqualand, Cape Province), Angola, Namibia, Botswana

Type locality: Interior of South Africa.  
Reproductionoviparous 
TypesType: BMNH 1865.5.4.103 (and possibly additional specimens)
Holotype: PEM R16066 (formerly AMG 6458) [frater] 
Diagnosis 
CommentP. formosus is listed by KLUGE (1993) as a valid species with question mark, but it is listed as subspecies of rugosus by RÖSLER 2000. LAMB & BAUER (2000) elevated P. rugosus barnardi and P. rugosus formosus to full species status. Distribution not yet corrected for elevation of barnardi and formosus to species status. 
References
  • Auerbach, R.D. 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Botswana, 295 pp.
  • Barts, M. 2010. Die Dickfingergeckos des südlichen Afrikas. Teil XI: Zur Haltung und Vermehrung von Pachydactylus rugosus SMITH, 1849. Sauria 32 (1): 3-12 - get paper here
  • Barts, M. 2021. Geckos aus dem südlichen Afrika. DATZ 2021 (5): 24-34
  • Barts, Mirko & Stefan Ballandat 2018. Vom Ugab nach Gaias – ein Abenteuer im nördlichen Namibia. Draco 16 (64): 36-44 [2015] - get paper here
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bauer, A.M. 2013. Geckos - The Animal Answer Guide. Johns Hopkins University Press, 159 pp.
  • Bauer, A.M., and Branch, W.R. 2003. The herpetofauna of the Richtersveld National Park, Northern Cape Province, Republic of South Africa. Herpetological Natural History 8: 111-160 [2001]
  • Bauer, Aaron M., Luis M. P. Ceríaco, Matthew P. Heinicke and David C. Blackburn. 2015. Geographic Distribution. Pachydactylus barnardi FitzSimons, 1941, Barnard's Rough Gecko. African Herp News (62): 35-37 - get paper here
  • Bauer, Aaron M.; Branch, William R. & Haacke, Wulf D. 1993. The herpetofauna of the Kamanjab area and adjacent Damaraland, Namibia. Madoqua (Windhoek) 18 (2): 117-145. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Branch, William R.; Pedro Vaz Pinto, Ninda Baptista, and Werner Conradie 2019. The Reptiles of Angola: History, Diversity, Endemism and Hotspots. Chapter 13 in: B. J. Huntley et al. (eds.), Biodiversity of Angola. Springer Verlag, pp. 283-334 - get paper here
  • Cimatti, E. 2007. Namibia - Introduction to a vast territory. Reptilia (GB) (50): 58-67 - get paper here
  • CONRADIE, WERNER; WILLIAM R. BRANCH, & GILLIAN WATSON 2019. Type specimens in the Port Elizabeth Museum, South Africa, including the historically important Albany Museum collection. Part 2: Reptiles (Squamata). Zootaxa 4576 (1): 001–045 - get paper here
  • Fitzsimons, V.F.M.; Brain, C.K. 1958. A Short account of the Reptiles of the Kalahari Gemsbok National Park. Koedoe, 1(1): 99-104 - get paper here
  • Fitzsimons,V.F.M. 1941. Descriptions of some new lizards from South Africa and a frog from southern Rhodesia. Annals Transvaal Mus. 20 (3): 273-281 - get paper here
  • Furman, Joe 1993. In search of the rough-scaled gecko, Pachydactylus rugosus. Dactylus 2 (1): 17-19
  • Gramentz, D. & Barts, M. 2004. Der Schrecklaut von Pachydactylus rugosus A. SMITH 1849. Sauria, Berlin 26 (1): 23-26 - get paper here
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Hewitt, J. 1927. Further descriptions of reptiles and batrachians from South Africa. Rec. Albany Mus. (Grahamstown) 3 (5): 371-415
  • Hewitt, J. 1935. Some new forms of batrachians and reptiles from South Africa. Rec. Albany Mus. 4: 283-357
  • Hewitt, John 1911. Records of South African Lacertilia and Amphibia, Addenda and Corrigenda. Annals Transvaal Museum 3 (1): 42-55. - get paper here
  • Lamb, T. & A. M. BAUER 2000. Relationships of the Pachydactylus rugosus group of geckos (Reptilia: Squamata: Gekkonidae). African Zoology 35 (1): 55-67 - get paper here
  • Lobón-Rovira J, Vaz Pinto P, S. Becker F, Tolley KA, Measey J, Bennet B, Boon B, de Sá S, Conradie W 2022. An updated herpetofaunal species inventory of Iona National Park in southwestern Angola. Check List 18(2): 289-321 - get paper here
  • Loveridge, A. 1947. Revision of the African lizards of the family Gekkondiae. Bull. Mus. Comp. Zool. Harvard 98: 1-469 - get paper here
  • McLachlan G R 1979. The taxonomy of Pachydactylus rugosus. J. Herp. Assoc. Africa (No. 21): 4-7 - get paper here
  • Mertens,R. 1955. Die Amphibien und Reptilien Südwestafrikas. Aus den Ergebnissen einer im Jahre 1952 ausgeführten Reise. Abh. senckenb. naturf. Ges. (Frankfurt) 490: 1-172 - get paper here
  • Pianka, Eric R. 1971. Lizard species density in the Kalahari Desert. Ecology 52 (6): 1024-1029 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Schleicher, Alfred 2015. Reptilien Namibias. Namibia Scientific  Society, 276 pp.
  • Schleicher, Alfred 2020. Reptiles of Namibia. Kuiseb Publishers, Windhoek, Namibia, 271 pp.
  • Smith, A. 1849. Illustrations of the Zoology of South Africa. 3 (Reptiles). Smith, Elder, and Co., London [facsimile published by Winchester Press, Johannesburg, 1977] - get paper here
  • Telford, N. S., ALEXANDER, G. J., BECKER, F. S., CONRADIE, W., JORDAAN, A., KEMP, L., ... & TOLLEY, K. A. 2022. EXTENSIONS TO THE KNOWN GEOGRAPHIC DISTRIBUTIONS OF REPTILES IN THE GREAT KAROO, SOUTH AFRICA. Herpetological Conservation and Biology, 17(1), 145-154 - get paper here
 
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