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Phoxophrys tuberculata HUBRECHT, 1881

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Hubrecht's Eyebrow Lizard 
SynonymPhoxophrys tuberculata HUBRECHT 1881: 51
Phoxophrys tuberculata — BOULENGER 1885: 280
Phoxophrys tuberculata — DE ROOIJ 1915: 94
Japalura robinsoni BOULENGER 1920: 287 (fide INGER 1960)
Phoxophrys tuberculata — INGER 1960: 225
Phoxophrys tuberculata — WERMUTH 1967: 75
Phoxophrys tuberculata — MANTHEY & GROSSMANN 1997: 197
Phoxophrys tuberculata — TEYNIÉ et al. 2010
Phoxophrys tuberculata — HARVEY et al. 2019 
DistributionIndonesia (W Sumatra: Jambi, Sumatera Barat, and Sumatera Utara Provinces)

Type locality: Batang, Singalang, West Sumatra.  
TypesType: BMNH 1946.8.13.92 (and possibly additional specimens).
Holotype: BMNH [robinsoni] 
DiagnosisDiagnosis (genus). Small draconine agamids reaching a SVL of at least 43 mm and distinguished from all other Agamidae by the following combination of characters: (1) tympanum and external auditory meatus absent, extrastapes attaching to skin; (2) head robust, 77.4–92.1% as wide as long, comprising 27.4–28.5% of SVL; (3) rostral single; (4) prominent tubercular scale positioned below corner of jaw; (5) supercilium projecting upward and outward as pointed, triangular flap; (6) dorsal scales heterogenous with larger tubercular scales interspersed among smaller keeled scales, 56–67 scales around midbody; (7) premaxillary processes of mandible abutting premaxilla; (8) frontal and parietal extensively sculptured, not containing pineal foramen; (9) transverse shorting of prefrontal bone allowing nasal–frontal contact; (10) squamosal extending anteriorly below jugal to curve ventrally and brace against vertical posterior margin of jugal; (11) angular process of mandible absent (Harvey et al. 2019: 94). 
CommentType species: Phoxophrys tuberculata HUBRECHT 1881 is the type species of the genus Phoxophrys HUBRECHT 1881. 
EtymologyHubrecht (1881) did not discuss the derivation of the name Phoxophrys. This feminine Latin noun in the nominative singular contains the Greek prefix phoxos meaning pointed or peaked and noun ophrys meaning eyebrow. The name likely refers to the arched, triangular supercilium of Ph. tuberculata and inspired coining of the common English name ‘‘eyebrow lizard,’’ as frequently used by various websites and in some recent treatments of the Agamidae (Murphy and Hanken 2018, Harvey et al. 2019). 
  • Aguilar-López JL, Luría-Manzano R, Pineda E, Canseco-Márquez L 2021. Selva Zoque, Mexico: an important Mesoamerican tropical region for reptile species diversity and conservation. ZooKeys 1054: 127-153 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, G.A. 1920. Results of an expedition to Korinchi Peak 12.400 ft, Sumatra. III. Reptiles & batrachians collected in Korinchi, West Sumatra by Messrs. H.C. Robinson & C. Boden Kloss. J. Federated Malay State Museum, Singapore, 8: 285-296 (other volumes in BHL)
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Denzer, W. 1996. Reptilien und Amphibien der Santubong Halbinsel, Sarawak, Borneo. Sauria 18 (4): 35-42 - get paper here
  • Harvey, Michael B., Larson, Thorton R., Jacobs, Justin L., Shaney, Kyle, Streicher, Jeffrey W., Hamidy, Amir, Kurniawan, Nia & Smit, Eric N. 2019. Phoxophrys After 60 Years: Review of Morphology, Phylogeny, Status of Pelturagonia, and a New Species from Southeastern Kalimantan. Herpetological Monographs 33 (1): 71-107 - get paper here
  • Hubrecht, A.A.W. 1881. On a new genus and species of Agamidae from Sumatra. Notes from the Leyden Museum 3: 51-52. - get paper here
  • Inger, R. F. 1960. A review of the agamid lizards of the genus Phoxophrys HUBRECHT. Copeia 1960 (3): 221 -225. - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
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