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Phrynocephalus versicolor STRAUCH, 1876

IUCN Red List - Phrynocephalus versicolor - Least Concern, LC

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
SubspeciesPhrynocephalus versicolor hispidus BEDRIAGA 1909
Phrynocephalus versicolor doriai BEDRIAGA 1909
Phrynocephalus versicolor kulagini BEDRIAGA 1909
Phrynocephalus versicolor versicolor STRAUCH 1876 
Common NamesE: Tuvan Toad-headed Agama
Russian: Пестрая круглоголовка
Chinese: 变色沙蜥 
SynonymPhrynocephalus versicolor STRAUCH 1876: 18
Phrynocephalus versicolor — BOULENGER 1885: 374
Phrynocephalus guentheri BEDRIAGA 1906: 168
Phrynocephalus versicolor var. Siebenrocki BEDRIAGA 1909
Phrynocephalus blanfordi BEDRIAGA 1907
Phrynocephalus ciliaris BEDRIAGA 1907
Phrynocephalus versicolor var. hispida BEDRIAGA 1909: 331
Phrynocephalus versicolor var. Doriai BEDRIAGA 1909: 332
Phrynocephalus versicolor var. Siebenrocki BEDRIAGA 1909: 333
Phrynocephalus versicolor var. Bogdanowi BEDRIAGA 1909: 334
Phrynocephalus helioscopus var. levis KASHTCHENKO 1909: 122
Phrynocephalus versicolor var. siebenrocki VON BEDRIAGA 1909: 232
Phrynocephalus Potanini BEDRIAGA 1909: 389
Phrynocephalus pewzowi
Phrynocephalus versicolor versicolor — NIKOLSKY 1915: 188
Phrynocephalus versicolor hispidus — NIKOLSKY 1915: 189
Phrynocephalus versicolor doriai — NIKOLSKY 1915: 190
Phrynocephalus versicolor siebenrocki — NIKOLSKY 1915: 191
Phrynocephalus versicolor bogdanowi — NIKOLSKY 1915: 191
Phrynocephalus rostralis ZAREVSKY 1930
Phrynocephalus parvulus ZAREVSKY 1927 (fide POPE 1935)
Phrynocephalus stejnegeri BANNIKOV 1958: 78 (nomen nudum)
Phrynocephalus versicolor — WERMUTH 1967: 89
Phrynocephalus immaculatus ZHAO 1995: 47
Phrynocephalus versicolor — WANG et al. 2003
Phrynocephalus (Phrynocephalus) versicolor versicolor — BARABANOV & ANANJEVA 2007
Phrynocephalus immaculatus — ANANJEVA 2011

Phrynocephalus versicolor hispidus BEDRIAGA 1909
Phrynocephalus versicolor hispida — TUNIYEV et al. 2014

Phrynocephalus versicolor kulagini BEDRIAGA 1909
Phrynocephalus versicolor var. Kulagini BEDRIAGA 1909: 329
Phrynocephalus versicolor kulagini — NIKOLSKY 1915: 189
Phrynocephalus (Phrynocephalus) versicolor kulagini — BARABANOV & ANANJEVA 2007

Phrynocephalus versicolor siebenrocki BEDRIAGA 1909
Phrynocephalus versicolor siebenrocki — MILTO & BARABANOV 2012: 165 
DistributionW China (Xinjiang, Gansu, Ningxia, Nei Mongol = Inner Mongolia), (Siberia), SE Russia (Tuva), Mongolia

doriai: Chian (Zinjiang); Terra typica restricta: "See Ebi-nor" [= Ebinur Lake, Xinjiang Uygur Autonomous Region, China].

hispida: Chian (Zinjiang); Terra typica restricta: "Dschungaria orientalis" [= Junggar Basin, Xinjiang Uygur Autonomous Region, China].

kalugini: NW Mongolia and Tyva Autonomous Region, Russia. Terra typica restricta: Chirgis-nur Lake in northwestern Mongolia.

Type locality: Alashan Desert, N China  
TypesLectotype: ZISP 3929.1 (designated by Peters, 1984: 36),
Lectotype: ZISP 5117.1 (designated by Peters, 1984). “See Ebinor” [Ebinur lake, Xinjiang Uygur Autonomous Region, China]. Leg: N. M. Przewalsky, 1878; however, Barabanov & Ananjeva 2007: 20 state that there is no type [doriai]
Lectotype: ZISP 5840, designated by Peters, 1984), “Sabul bei Kirgis-Nor” [Chirgis-nur lake, Uvs province, Mongolia]. Leg: G. N. Potanin, 1889 [kulagini]
Holotype: ZISP 5156, “zwischen Metschin-Ola und der Stadt Barkul” [between Mieh-Ch’eng-O-La Shan (about 30 km northeast of Barkol) and Barkol, approx. 43°55’ N 93°00 E, Barkol county, Xinjiang Uygur Autonomous Region, China]. Leg: G. N. Potanin, 1878 [siebenrocki]
Lectotype: ZISP 6637.1 (designated by Peters, 1984: 37) [hispidus] 
CommentGolubev (1993) gives a description of P. v. hispida and P. v. doriai. Oviparous. Wang & Fu (2004) suggested to apply the name P. versicolor only to the populations in the northwestern part of its distribution area. Populations from Tengger Desert, Ordos highland, and further east include P. frontalis, P. przewalskii, and P. versicolor and should be called P. przewalskii. Some populations may contain a new species that remains to be described.

Guo & Wang (2007) recognized populations in Kuytun, Xinjiang Uygur Autonomous Region as P. guttatus instead of P. versicolor.

Synonymy after BARABANOV & ANANJEVA (2007) who consider doriai and hispidus as synonyms of versicolor (and thus recognize only 2 subspecies: versicolor and kulagini). 
  • ANANJEVA, Natalia B.; Xianguang GUO and Yuezhao WANG 2011. Taxonomic Diversity of Agamid Lizards (Reptilia, Sauria, Acrodonta, Agamidae) from China: A Comparative Analysis. Asian Herpetological Research 2 (3): 117-128 - get paper here
  • Ananjeva, Natalia B.;Tuniyev, Boris S. 1992. Historical biogeography of the Phrynocephalus species of the USSR. Asiatic Herpetological Research 4: 76-98 - get paper here
  • BARABANOV, A.V. & N.B. ANANJEVA 2007. Catalogue of the available scientific species-group names for lizards of the genus Phrynocephalus Kaup, 1825 (Reptilia, Sauria, Agamidae). Zootaxa 1399: 1-56 - get paper here
  • Bedriaga, J. von 1907. Wissenschaftliche Resultate der von N. M. Przewalski nach Central-Asien unternommenen Reisen.Zoologischer Theil, Amphibien und Reptilien, Band 3, Abt. 1, Lieferung 2: Lacertilia (Gekkonidae, Agamidae). Kaiserl. Akad. Wissensch., St. Petersburg, pages 71-278
  • Bedriaga, J.V. 1909. Amphibien und Reptilien. In: Wissenschaftliche Resultate der Reisen N.M. Przewalskijs durch Zentralasien. Zoologische Teil. Band 3. Part 1. Lacertilia. (in Russian). Sankt-Petersburg, Vol 3 (1) 3: 73-102, 367
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
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  • Golubev, M.L. 1993. The variegated toad agama in Djungar Gate (Eastern Kazakstan) with notes on certain systematic problems of Phrynocephalus versicolor Str. (Reptilia, Agamidae). Asiatic Herpetological Research 5: 51-58 - get paper here
  • Guo, X. & Wang, Y. 2007. Partitioned Bayesian analyses, dispersal–vicariance analysis, and the biogeography of Chinese toad-headed lizards (Agamidae: Phrynocephalus): A re-evaluation. Molecular Phylogenetics and Evolution <br />45 (2): 643-662 - get paper here
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  • Milto, Konstantin D.; Andrei V. Barabanov 2012. A Catalogue of the Agamid and Chamaeleonid Types in the Collection of the Zoological Institute, Russian Academy of Sciences, St. Petersburg. Russ. J. Herpetol. 19 (2): 155-170 - get paper here
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  • Pang, Junfeng; Yuezhao Wang, Yang Zhong, A. Rus Hoelzel, Theodore J. Papenfuss, Xiaomao Zeng, Natalia B. Ananjeva and Ya-ping Zhang 2003. A phylogeny of Chinese species in the genus Phrynocephalus (Agamidae) inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 27: 398-409 - get paper here
  • Pope, CLIFFORD H. 1935. The Reptiles of China. Turtes, Crocodilians, Snakes, Lizards. Amer. Mus. Nat. Hist., New York, Nat. Hist. Central Asia, 10: lii, 1-604 - get paper here
  • Strauch in: PRZEWALSKI, N.M. 1876. Mongoliya i Strana Tangutov. Tryokhletneye puteshestviye v Vostochnoj Nagoruoj Asii (= Mongolia and the Tangut Country. A Three-Year Travel in Eastern High Asia). Volumes 1-2. Imper. Russ. Geogr. Soc., St. Petersburg (in Russian; English edition, London 1876, French edition, Paris 1880, German edition, Jena 1887; abridged Russian edition with added notes, Moscow 1946). - get paper here
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp. - get paper here
  • TANG, Xiaolong; Feng YUE, Ming MA, Ningbo WANG, Jianzheng HE and Qiang CHEN 2012. Effects of Thermal and Hydric Conditions on Egg Incubation and Hatchling Phenotypes in Two Phrynocephalus Lizards. Asian Herpetological Research 3 (3): 184–191 - get paper here
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