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Phymaturus nevadoi CEI & ROIG, 1975

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Higher TaxaLiolaemidae, Iguania, Sauria, Squamata (lizards)
Common Names 
SynonymPhymaturus patagonicus nevadoi CEI & ROIG 1975
Phymaturus nevadoi CEI & CASTRO 1975
Phymaturus nevadoi — LOBO & QUINTEROS 2005
Phymaturus nevadoi — LOBO et al. 2021 
DistributionArgentina (Mendoza)

Type locality: Argentina, Mendoza, Agua de la India, Muerta, Nevado Mts., 1750 m [35° 46’ S, 68° 27’ W]  
TypesHolotype: IBAUNC 999-1 
DiagnosisComparisons: for a table of characters across the payuniae clade (including nevadoi), see Lobo et al. 2021: Table 2.

Coloration: The dorsal white spots are large (formed by 9–16 scales) in P. nevadoi, but small in P. niger sp. nov. (formed by no more than 6 scales; Table 2). Female P. nevadoi possess dorsal ocelli, which are absent in P. niger sp. nov. The throat pattern in P. nevadoi is sparse and formed by thin lines (Fig. 6C), while it is formed by a dense reticulation of thin, interconnected lines in P. niger sp. nov. (some individuals lack any pattern; Table 2). The ventral pattern of the tail is present in P. nevadoi but absent in P. niger sp. nov. The belly color is pink/orange in male P. nevadoi and yellow in male P. niger sp. nov. (Table 2). Phymaturus nevadoi is slightly smaller (x̅ = 86.5 ± 4.16 mm SVL) than P. niger sp. nov. (x̅ = 91.0 ± 4.9 mm SVL). No P. nevadoi are completely melanic. Phymaturus nevadoi exhibits fewer scales around midbody (x̅ = 203.7 ± 9.9) than P. niger sp. nov. (x̅ = 221.1 ± 16.3)
Dorsal ocelli are absent in P. robustus sp. nov. but common in female P. nevadoi. The throat pattern in P. robustus sp. nov. (Fig. 6D) is sparce, formed by thick lines, whereas P. nevadoi’s throat pattern is scarce, but formed by thin lines (Fig. 6C; Table 2). Also, individuals of P. robustus sp. nov. display a conspicuous black line behind the mental scale on the midline (Fig. 6D), absent in P. nevadoi (Table 2). The dorsal tail pattern of P. robustus sp. nov. is either variegated or absent, different from the spotted pattern of P. nevadoi (Table 2). The ventral pattern of the tail is absent in P. robustus sp. nov., different from that of P. nevadoi. Phymaturus robustus sp. nov. exhibits a higher number of scales around midbody than P. nevadoi (x̅ = 222.2 ± 13.1 and x̅ = 203.7 ± 9.9, respectively). Phymaturus robustus sp. nov. is larger than P. nevadoi (x̅ = 96.9 ± 4.8 mm SVL and x̅= 86.5 ± 4.2 mm SVL,respectively). The sulcus spermaticus is open in the hemipenes of P. robustus sp. nov. but closed in P. nevadoi. The sulcus bifurcation is distal in P. robustus sp. nov., unlike in P. nevadoi (Table 2) (Lobo et al. 2021). 
CommentFull species fide ETHERIDGE 1995.

Diet: herbivorous

This species belongs to the patagonicus group (fide LOBO et al. 2010). 
EtymologyNamed after the type locality. 
  • AVILA, LUCIANO JAVIER; CRISTIAN HERNAN FULVIO PEREZ, DANIEL ROBERTO PEREZ & MARIANA MORANDO 2011. Two new mountain lizard species of the Phymaturus genus (Squamata: Iguania) from northwestern Patagonia, Argentina. Zootaxa 2924: 1–21 - get paper here
  • Avila, Luciano Javier; Cristian Hernan Fulvio Perez, IGNACIO MINOLI & Mariana Morando 2014. A new lizard of the Phymaturus genus (Squamata: Liolaemidae) from Sierra Grande, northeastern Patagonia, Argentina. Zootaxa 3793 (1): 099–118 - get paper here
  • AVILA, LUCIANO JAVIER; LORENA ELIZABETH MARTINEZ & MARIANA MORANDO 2013. Checklist of lizards and amphisbaenians of Argentina: an update. Zootaxa 3616 (3): 201–238 - get paper here
  • Cei, J.M. & ROIG, V.G. 1975. A new lizard from the Sierra del Nevado Mountains,Central Argentina. Journal of Herpetology 9 (2): 256 - get paper here
  • Corbalán, Valeria; Guillermo Deband , José Alejandro Scolaro , and Agustina Ojeda 2016. DNA Barcoding of Phymaturus Lizards Reveals Conflicts in Species Delimitation within the Patagonicus Clade. Journal of Herpetology 50 (4): 654-666 - get paper here
  • GONZÁLEZ-MARÍN, ANDREA; CRISTIAN hERNÁN FULVIO PÉREZ, IGNACIO MINOLI, Mariana Morando, Luciano Javier Avila 2016. A new lizard species of the Phymaturus patagonicus group (Squamata: Liolaemini) from northern Patagonia, Neuquén, Argentina. Zootaxa 4121 (4): 412–430 - get paper here
  • Lobo, F. and S. Quinteros 2005. Taxonomic studies of the genus Phymaturus (Iguania: Liolaemidae): Redescription of Phymaturus patagonicus Koslowsky 1898, and revalidation and redescription of Phymaturus spurcus Barbour 1921. Journal of Herpetology 39 (4): 533-540 - get paper here
  • Lobo, F., Barrasso, D. A., Hibbard, T., Quipildor, M., Slodki, D., Valdecantos, S., & Basso, N. G. 2021. Morphological and genetic divergence within the Phymaturus payuniae clade (Iguania: Liolaemidae), with the description of two new species. South American Journal of Herpetology 20 (1): 42-66 - get paper here
  • Lobo, Fernando; Cristian Abdala and Soledad Valdecantos 2010. Taxonomic Studies of the Genus Phymaturus (Iguania: Liolaemidae): Description of Four New Species. South American J. Herp. 5 (2):102-126 - get paper here
  • LOBO, FERNANDO; CRISTIAN ABDALA & SOLEDAD VALDECANTOS 2012. Morphological diversity and phylogenetic relationships within a South-American clade of iguanian lizards (Liolaemidae: Phymaturus). Zootaxa 3315: 1–41 - get paper here
  • Scolaro, J.A. & Pincheira-Donoso, D. 2010. Lizards at the end of the world: Two new species of Phymaturus of the patagonicus clade (Squamata, Liolaemidae) revealed in southern Patagonia of Argentina. Zootaxa 2393: 17–32 - get paper here
  • SCOLARO, JOSÉ ALEJANDRO; FAUSTO MÉNDEZ DE LA CRUZ & NORA R. IBARGÜENGOYTÍA 2012. A new species of Phymaturus of the patagonicus clade (Squamata, Liolaemidae) from isolated plateau of southwestern Rio Negro Province, Argentina. Zootaxa 3451: 17–30 - get paper here
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