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Platyceps florulentus (GEOFFROY SAINT-HILAIRE, 1827)

IUCN Red List - Platyceps florulentus - Least Concern, LC
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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
SubspeciesPlatyceps florulentus florulentus (GEOFFROY SAINT-HILAIRE 1827)
Platyceps florulentus keniensis (PARKER 1932)
Platyceps florulentus perreti (SCHÄTTI 1988) 
Common NamesE: Flowered Racer, Geoffroys’ Racer
G: Ägyptische Zornnatter 
SynonymColuber florulentus GEOFFROY SAINT-HILAIRE 1827: 146, 151
Zamenis florulentus — DUMÉRIL, BIBRON & DUMÉRIL 1854: 693
Zamenis florulentus — BOULENGER 1893: 402
Zamenis florulentus — CALABRESI 1925: 106
Zamenis florulentus — SCORTECCI 1931: 17
Coluber florulentus — LOVERIDGE 1936
Coluber florulentus florulentus — LOVERIDGE 1955
Coluber florulentus —SCHÄTTI & WILSON 1986
Haemorrhois florulentus — MEIRTE 1992
Coluber florulentus — LARGEN & RASMUSSEN 1993
Haemorrhois florulentus — LEBRETON 1999
Platyceps florulentus — NAGY et al. 2004
Platyceps florulentus — SCHLÜTER 2005
Platyceps florulentus — WALLACH et al. 2014: 564
Platyceps florulentus — SPAWLS et al. 2018: 478

Platyceps florulentus keniensis (PARKER 1932)
Coluber keniensis PARKER 1932: 220
Coluber keniensis — SCHÄTTI & WILSON 1986
Coluber florulentus keniensis — SCHÄTTI 1988: 98
Haemorrhois florulentus keniensis — MEIRTE 1992

Platyceps florulentus perreti (SCHÄTTI 1988)
Coluber florulentus perreti SCHÄTTI 1988: 100
Haemorrhois florulentus perreti — CHIRIO & LEBRETON 2007
Platyceps florulentus perreti — TRAPE & BALDÉ 2014: 323
Platyceps perreti — SCHÄTTI 2014: 384 
DistributionEgypt (North Sinai [HR 32: 59]), Sudan (Jumhūriyyat), Republic of South Sudan (RSS), Ethiopia, Eritrea, Somalia, Kenya, Uganda, Cameroon

keniensis: Type locality: Lake Baringo

perreti: Cameroon, Nigeria; Type locality: “Soulede, Mokolo, N Kamerun”

Type locality: Egypt  
Reproductionoviparous 
TypesHolotype: lost (probably, fide SCHÄTTI 1988).
Holotype: MHNG 1464.45, male; paratypes: BMNH 1975.1074 [juv., probably from Bambour (Bambur) area, Nigeria], BMNH 1975.1141-42 (juv. female and ad. female, Bambour), USNM 125691 (female, Marama, Nigeria), ZFMK 18686 [juv. female, Benoue (Garoua), Kamerun]. [perreti]
Holotype: BMNH 1946.1.13.87 (orig. 1932.5.2.7), collected by E.B. Worthington, 1930 [keniensis] 
DiagnosisDiagnosis (perreti). A subspecies of C. florulentus distinguished from the nominate race by 25 (instead of 21-23) dorsal scale rows at midbody. It differs from f. florulentus and f. keniensis in higher ventral counts (219 versus 214 or fewer in males, 226-231 against 205-228 in females), and more maxillary teeth (16-17 instead of 15 or less). 
CommentDistribution: Not explicitely listed by SCHLEICH, KÄSTLE & KABISCH 1996. The author is “GEOFFROY 1809: 67” fide DUMÉRIL et al. 1854. Reports from Libya represent Hemorrhois algirus or possibly P. rogersi (Bauer et al. 2017). For a map see Sindaco et al. 2013.

Subspecies: Schätti 2014: 384 elevated perreti to full species. However, the only difference he mentions is that perreti has 25 dorsals (vs. 27-29 in the Platyceps florulentus complex) 
References
  • Bauer, Aaron M.; Jonathan C. DeBoer , Dylan J. Taylor 2017. Atlas of the Reptiles of Libya. Proc. Cal. Acad. Sci. 64 (8): 155-318 - get paper here
  • Bons, J. 1962. Notes sur trois couleuvres africaines: Coluber algirus, Coluber florulentus et Coluber hippocrepis. Description de Coluber algirus villiersi subsp. nov. Bull. Soc. Sci. nat. Maroc 42: 61-86.
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Broadley, D.G. 1994. A collection of snakes from eastern Sudan, with the description of a new species of Telescopus Wagler, 1830 (Reptilia: Ophidia). Journal of African Zoology 108 (2): 201-208.
  • Broadley,D.G. & Schätti,B. 2000. A new species of Coluber from northern Namibia (Reptilia: Serpentes). Madoqua 19 (2): 171-174 [1997] - get paper here
  • Calabresi, Enrica 1925. Anfibi e rettili raccolti dal Signor Ugo Ignesti nell'Abissinia settentrionale. Atti della Società Italiana di Scienze Naturali, e del Museo Civico di Storia Naturale, Milano 64: 100-109 - get paper here
  • Chippaux, Jean-Philippe & Kate Jackson 2019. Snakes of Central and Western Africa. Johns Hopkins University Press, 448 pp. [detaileld review in HR 51 (1): 161] - get paper here
  • Geniez, Philippe 2018. Snakes of Europe, North Africa and the Middle East. Princeton University Press, 384 pp. - get paper here
  • Geoffroy Saint-Hilaire, E. 1827. Description des Reptiles. In: Savigny, M. J. C. L. de, Description de l’Égypte ou recueil des observations et des recherches Quintana Roo ont été faites en Égypte pendant l’expedition de l’Armée Française, publié par les ordres de sa Majesté l’Empere Paris, L’imprimerie impériale. pp. 121-160 (115-184) [1809?] - get paper here
  • Gruber, U. 2009. Die Schlangen Europas, 2. Aufl. Kosmos Naturführer, 266 pp.
  • Kirchhof S, Wasonga V, Mazuch T, Spawls S, Malonza KP. 2023. An annotated checklist of the herpetofauna of the Sibiloi National Park in northern Kenya based on field surveys. Amphibian & Reptile Conservation 17(1/2) [General Section]: 1–18 (e324) - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Largen,M.J. & Rasmussen,J.B. 1993. Catalogue of the snakes of Ethiopia (Reptilia Serpentes), including identification keys. Tropical Zoology 6: 313-434 - get paper here
  • LeBreton, Matthew 1999. A working checklist of the herpetofauna of Cameroon. Netherlands Committee for IUCN, 160 pp.
  • Loveridge, A. 1936. African reptiles and amphibians in the Field Museum of Natural History. Zool. Ser. Field Mus. Nat. Hist., Chicago, 22 (1): 1-122 - get paper here
  • Loveridge, A. 1956. On snakes collected in the Anglo-Egyptian Sudan by J.S. Owen, Esq. Sudan Notes Rec. 36: 37-56 [1955]
  • Meirte, D. 1992. Clés de détermination des serpents d'Afrique. Museum Royal d'Afrique Centrale, Tervuren Belgique Annual Series Octavo Science Zoologique 267:1-152
  • Nagy, Z.T.; R. Lawson, U. Joger, M. Wink 2004. Molecular systematics of racers, whipsnakes and relatives (Reptilia: Colubridae) using mitochondrial and nuclear markers. Journal of Zoological Systematics and Evolutionary Research 42 (3): 223-233 - get paper here
  • Osman, H.E.S.I., & Sir, N.T.E. 1988. The snakes of the Sudan. 1. The snakes of Khartoum Province. The Snake 20 (1): 74-79.
  • Parker,H.W. 1932. Scientific results of the Cambridge expedition to the east African lakes, 1930-31. 5. Reptiles and amphibians. Zool. J. Linnean Soc. 38: 213-229 - get paper here
  • Pitman,C.R.S. 1974. A guide to the snakes of Uganda. Codicote, Wheldon & Wesley, L., 290 pp.
  • Schätti B; Wilson L D 1986. Coluber Linnaeus. Holarctic racers. Catalogue of American Amphibians and Reptiles ( 399: 1-4 - get paper here
  • Schätti, B. 1988. Systematics and phylogenetic relationships of Coluber florulentus Geoffroy, 1827 (Reptilia, Serpentes). Tropical Zoology 1 (1): 95-116 - get paper here
  • Schätti, Beat; Frank Tillack & Christoph Kucharzewski 2014. Platyceps rhodorachis (Jan, 1863) – a study of the racer genus Platyceps Blyth, 1860 east of the Tigris (Reptilia: Squamata: Colubridae). Vertebrate Zoology 64 (3): 297-405 - get paper here
  • Schlüter, U. 2005. Die Zornnattern Nordafrikas. Reptilia (Münster) 10 (52): 56-64 - get paper here
  • Scortecci, Giuseppe 1931. Contributo alla conoscenza dei rettili e degli anfibi della Somalia, dell'Eritrea e dell'Abissinia. Boll. Mus. Zool. Univ. Torino, (ser. 3) 41 (10): 1-26 [1930]
  • Sindaco, R.; Alberto Venchi & Cristina Grieco 2013. The Reptiles of the Western Palearctic, Volume 2: Annotated Checklist and Distributional Atlas of the Snakes of Europe, North Africa, Middle East and Central Asia, with an Update to Volume 1. Edizioni Belvedere, Latina (Italy), 543 pp. - get paper here
  • Sochurek,E. 1979. Die Schlangen Nordafrikas. Mitt. Zool. Ges. Braunau 3 (8/9): 219-226
  • Spawls, Stephen; Tomáš Mazuch& Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • TRAPE, JEAN-FRANÇOIS & CELLOU BALDÉ 2014. A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa 3900 (3): 301–338 - get paper here
  • Venchi, Alberto and Roberto Sindaco 2006. Annotated checklist of the reptiles of the Mediterranean countries, with keys to species identification. Part 2 -Snakes (Reptilia, Serpentes). Annali del Museo Civico di Storia Naturale "G. Doria", Genova, XCVIII: 259-364
  • Wall, F. 1908. Extension of the habitat of the Snake Zamenis florulentus. J. Bombay nat. Hist. Soc. 18: 689-690 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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