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Podarcis liolepis (BOULENGER, 1905)

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Higher TaxaLacertidae, Lacertinae, Lacertini, Sauria, Lacertoidea, Squamata (lizards)
Common NamesE: Catalonian Wall Lizard, Columbretes Wall Lizard
G: Columbretes Mauereidechse
D: Columbretes Muurhagedis 
SynonymLacerta muralis var. liolepis BOULENGER 1905
Podarcis hispanica atrata (BOSCÁ, 1916)
Lacerta atrata columbretensis EISENTRAUT 1930
Lacerta atrata mancolibrensis EISENTRAUT 1930
Lacerta bocagei liolepis — MERTENS & MÜLLER 1940
Lacerta bocagei columbretensis — EISENTRAUT 1949: 57
Lacerta bocagei atrata — EISENTRAUT 1949: 130
Lacerta bocagei liolepis — EISENTRAUT 1949: 57
Podarcis hispanica atrata — SALVADOR 1986: 77
Podarcis hispanica atrata — BAUER & GÜNTHER 1995
Podarcis hispanica atrata — PÉREZ-MELLADO in SALVADOR 1998
Podarcis atrata — CASTILLA et al. 1998
Podarcis atrata — OLIVERIO et al. 2000
Podarcis atrata — ARNOLD et al. 2007
Podarcis atratus — SINDACO & JEREMČENKO 2008
Podarcis liolepis — RENOULT et al. 2010
Podarcis liolepis — SPEYBROECK et al. 2020 
DistributionSE Spain (Columbretes Island, off Castellón), Andorra, France

Type locality: Valencia

atrata: Type locality: Les Ferreres/Columbreten (Spain)

mancolibrensis: Type locality: Mancolibre/Columbreten

columbretensis: Type locality: Columbrete Grande  
TypesSyntypes: BMNH 1946.9.1.33-35 
CommentType not in BMNH fide P. Campbell, pers. comm., 9 July 2014.

Synonymy: Mayer listed atrata under liolepis. Castilla et al. (1998) elevated this form to full species status but few authors seem to have accepted this decision.

Distribution: see map in Caeiro-Dias et al. 2018. Börner (2019) discusses possibly introduced populations in Germany. 
  • Arnold, E.N.; Arribas, O. & Carranza, S. 2007. Systematics of the Palaearctic and Oriental lizard tribe Lacertini (Squamata: Lacertidae: Lacertinae), with descriptions of eight new genera. Zootaxa 1430: 1-86 - get paper here
  • Bauer, Aaron M.;Günther, Rainer 1995. An annotated type catalogue of the lacertids (Reptilia: Lacertidae) in the Zoological Museum, Berlin. Mitt. Zool. Mus. Berlin 71 (1): 37-62 - get paper here
  • Berroneau, M. et al. 2010. Guide des Amphibiens et Reptiles d’Aquitaine. Association Cistude Nature, 180 pp.
  • Börner, Achim-Rüdiger 2019. Herbstbesuch bei den Lacerten auf Burg Hardenberg bei Göttingen. Saurologica (6): 68 pp., 65 figs [published Köln 12. XII. 2019] - get paper here
  • Boscá, E. 1916. Dos observaciones a propósito de la Lacerta muralis en España. Bol. R. Soc. Española Hist. Nat. 16: 327-330. - get paper here
  • Boulenger, G. A. 1905. A contribution to the knowledge of the varieties of the Wall-Lizard (Lacerta muralis) in western Europe and North Africa. Transact. zool. Soc. London, 17: 351-420 - get paper here
  • Caeiro-Dias, Guilherme, Carla Luís, Catarina Pinho, Pierre-André Crochet, Neftalí Sillero and Antigoni Kaliontzopoulou 2018. Lack of congruence of genetic and niche divergence in Podarcis hispanicus complex. J Zool Syst - get paper here
  • Carneiro, D., E. García-Muñoz, A. Kaliontzopoulou, G. A. Llorente & M. A. Carretero 2015. Comparing ecophysiological traits in two Podarcis wall lizards with overlapping ranges. Salamandra 51 (4): 335-344 - get paper here
  • Castilla, Aurora M.;Fernandez-Pedrosa, Victoria; Harris, D. James; Gonzalez, Ana;Latorre, Amparo; Moya, Andrés 1998. Mitochondrial DNA divergence suggests that Podarcis hispanica atrata (Squamata: Lacertidae) from the Columbretes Islands merits specific dictinction. Copeia 1998 (4): 1037-1040 - get paper here
  • Dieckmann, Martin 2018. Das südliche Frankreich und seine Herpetofauna. Terraria-Elaphe 2018 (2): 14-28 - get paper here
  • Eisentraut, M. 1949. Die Eidechsen der Spanischen Mittelmeerinseln und ihre Rassenaufspaltung im Lichte der Evolution. Mitteilungen aus dem Zoologischen Museum in Berlin, 26: 5-225 - get paper here
  • Kaliontzopoulou, A., Carretero, M.A. & Llorente, G., 2012. Morphology of Podarcis wall lizards from the Iberian Peninsula and North Africa: patterns of variation in a putative cryptic species complex. Zoological Journal of the Linnean Society, 164: 173-193 - get paper here
  • Kaliontzopoulou, A., Pinho, C., Harris, D.J. & Carretero, M.A., 2011. When cryptic diversity blurs the picture: a cautionary tail from Iberian and North African Podarcis wall lizards. Biological Journal of the Linnean Society, 103: 779-800 - get paper here
  • Kwet, A. 2010. Reptilien und Amphibien Europas. Kosmos-Verlag, Stuttgart, 252 pp.
  • Kwet, Axel & Benny Trapp 2014. Liste der Reptilien Europas. Draco 15 (60): 72-79 - get paper here
  • Kwet, Axel & Benny Trapp 2014. Reptilien der griechischen Ägäisinseln. Draco 15 (60): 58-65 - get paper here
  • Mertens, R., and L. Müller. 1940. Liste der Amphibien und Reptilien Europas. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft. Frankfurt am Main. 451: 1-62 - get paper here
  • Oliverio, M.; Bologna, M.A. & Mariottini, P. 2000. Molecular biogeography of the Mediterranean lizards Podarcis Wagler, 1830 and Teira Gray, 1838 (Reptilia, Lacertidae). Journal of Biogeography 27: 1403–1420. - get paper here
  • Pérez-Mellado, V., Galindo, M.P. 1986. Sistemática de Podarcis (Sauria, Lacertidae) Ibéricas y Norteafricanas mediante Técnicas Multidimensionales. Salamanca, Sér. Manual. Univ. Salamanca.
  • Pottier G., Paumier J.-M., Tessier M., Barascud Y., Talhoët S., Liozon R., D’Andurain P., Vacher J.-P., Barthe L., Heaulmé V., Esslinger M., Arthur C.-P., Calvet A., Maurel C. & Redon H. 2008. Atlas de répartition des reptiles et amphibiens de Midi-Pyrénées. Les atlas naturalistes de Midi-Pyrénées. Nature Midi-Pyrénées, Toulouse, 126 pp.
  • RENOULT, JULIEN P.; PHILIPPE GENIEZ, PAUL BACQUET, CLAUDE P. GUILLAUME, & PIERREANDRE CROCHET 2010. Systematics of the Podarcis hispanicus-complex (Sauria, Lacertidae) II: the valid name of the north-eastern Spanish form. Zootaxa 2500: 58–68 - get paper here
  • Salvador, A. 1986. Podarcis hispanica - Iberische Mauereidechse. In: Böhme,W. (ed.), Handbuch der Reptilien und Amphibien Europas, Band 2/II., Echsen III (Podarcis). Aula-Verlag Wiesbaden, pp. 71-82
  • Schlüter, U. 2011. Die Lacertiden Andorras. Die Eidechse 22 (3): 83-92 - get paper here
  • Schulte, Ulrich; Gassert, Franz; Geniez, Philippe; Veith, Michael; Hochkirch, Axel 2012. Origin and genetic diversity of an introduced wall lizard population and its cryptic congener. Amphibia-Reptilia 33 (1): 129-140 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Speybroeck, Jeroen; Wouter Beukema, Christophe Dufresnes, Uwe Fritz, Daniel Jablonski, Petros Lymberakis, Iñigo Martínez-Solano, Edoardo Razzetti, Melita Vamberger, Miguel Vences, Judit Vörös0, Pierre-André Crochet 2020. Species list of the European herpetofauna – 2020 update by the Taxonomic Committee of the Societas Europaea Herpetologica.. Amphibia-Reptilia 41: 139-189 - get paper here
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