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Tenuidactylus longipes (NIKOLSKY, 1896)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
SubspeciesTenuidactylus longipes longipes (NIKOLSKY 1896)
Tenuidactylus longipes microlepis (LANTZ 1918) 
Common NamesE: Nikolsky's Long-Toed Gecko, Long-Legged Thin-Toed Gecko 
SynonymGymnodactylus longipes NIKOLSKY 1896
Cyrtodactylus longipes — ANDERSON 1963: 474
Tenuidactylus longipes — SZCZERBAK & GOLUBEV 1984: 53
Cyrtopodion (Tenuidactylus) longipes — BÖHME 1985: 96
Tenuidactylus longipes — KHAN & TASNIM 1990
Tenuidactylus longipes — KLUGE 1993
Cyrtopodion (Tenuidactylus) longipes — RÖSLER 1995: 113
Cyrtopodion longipes — ANDERSON 1999: 163
Tenuidactylus longipes — SZCZERBAK 2003
Indogekko longipes — KHAN 2003
Indogekko longipes — KRYSKO et al. 2007
Indogekko longipes — KHAN 2008
Cyrtopodion (Tenuidactylus) longipes — SINDACO & JEREMČENKO 2008
Tenuidactylus longipes — BAUER et al. 2013
Tenuidactylus longipes — RÖSLER 2018: 69

Tenuidactylus longipes longipes (NIKOLSKY 1896)
Gymnodactylus longipes longipes — GORELOV et al. 1974: 35
Tenuidactylus longipes longipes — SZCZERBAK & GOLUBEV 1986: 156
Cyrtopodion longipes longipes — ANDERSON 1999: 163
Cyrtopodion (Tenuidactylus) longipes longipes — RÖSLER 2000: 74

Tenuidactylus longipes microlepis (LANTZ 1918)
Gymnodactylus microlepis LANTZ 1918
Gymnodactylus fedtschenkoi — SMITH 1935: 41 (part.)
Gymnodactylus (Cyrtodactylus) longipes microlepis — GORELOV et al. 1974: 35
Tenuidactylus longipes microlepis — SZCZERBAK & GOLUBEV 1986: 156
Cyrtopodion longipes microlepis — ANDERSON 1999: 164
Cyrtopodion (Tenuidactylus) longipes microlepis — RÖSLER 2000: 74
Tenuidactylus microlepis — NAZAROV & POYARKOV 2013 
DistributionE Iran, Afghanistan, former USSR, S Turkmenistan, Afghanistan, S Uzbekistan

longipes: E Iran, adjacent Afghanistan

microlepis: SW Turkmenistan, Iran

Type locality: Tedzhen, S Turkmenistan  
TypesLectotype: ZISP (ZIL) 8810, designated by SZCZERBAK & GOLUBEV 1986:153; Paralectotypes (former syntypes): ZISP (ZIL) 8809 (6), 8810 (3), 8811
Lectotype: ZMMU 151 (designated by Szczerbak & Golubev, 1986) [microlepis]
Holotype: CAS 120322 [voraginosus] 
DiagnosisDiagnosis (longipes): Enlarged dorsal tubercles separated by small scales; 27-36 abdominal scales across middle ofbelly; 26-30 dorsal tubercles in paravertebral row from occiput to level of vent; 32-38 preanal and femoral pores (total both sides), series not separated by scales lacking pores, females with corresponding depressions in scales; 12-16 interorbital scales; length of forelimb 38-45% of snout-vent length; first pair of postmentals usually in relatively broad or point contact, or separated by one or two scales (Anderson 1999: 163, who listed this species under Cyrtopodion).

Diagnosis (microlepis): Dorsal tubercles separated by small scales; 31-33 tubercles in paravertebral row from occiput to level of groin; 35-39 abdominal scales across middle of beIIy; total of 27-35 preanal and femoral pores (seen as slight depressions in the scales in females), the series not separated by scales lacking pores; 12-14 interorbital scales (Anderson 1999: 164). 
CommentSynonymy partly after KHALIKOV & ANANJEVA (pers. comm.)

Subspecies: Cyrtopodion longipes voraginosus (LEVITON & ANDERSON 1984) is now considered as a valid species.

Distribution: See map in SMID et al. 2014 for distribution in Iran. 
EtymologyNamed after Latin longus = long + L. pes = leg, foot (Borror, 1988). 
  • ANANJEVA, NATALIA B.; KONSTANTIN D. MILTO, ANDREI V. BARABANOV, EVGENY A. GOLYNSKY 2020. An annotated type catalogue of amphibians and reptiles collected by Nikolay A. Zarudny in Iran and Middle Asia. Zootaxa 4722 (2): 101–128 - get paper here
  • Ananjeva, NB, Orlov NL; Khalikov RG; Darevsky IS; Ryabov IS & Barabanov AV 2006. The Reptiles of North Eurasia. Taxonomic Diversity, Distribution, Conservation Status [this comprises the territory of the former Soviet Union and Mongolia]. Pensoft Series Faunistica 47, 250 pp.
  • Anderson, S.C. 1963. Amphibians and Reptiles from Iran. Proc. Cal. Acad. Sci. Ser. 4, 31(16): 417-498 - get paper here
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • BAUER, AARON M.; RAFAQAT MASROOR, JAMES TITUS-MCQUILLAN, MATTHEW P. HEINICKE,, JUAN D. DAZA & TODD R. JACKMAN 2013. A preliminary phylogeny of the Palearctic naked-toed geckos (Reptilia: Squamata: Gekkonidae) with taxonomic implications. Zootaxa 3599 (4): 301–324 - get paper here
  • Böhme, W. 1985. Zur Nomenklatur der paläarktischen Bogenfingergeckos, Gattung Tenuidactylus Szczerbak & Golubev 1984 (Reptilia: Gekkonidae). Bonner Zoologische Beiträge 36: 95-98 - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Gorelov, Y.K., Darevsky, I.S. & Szczerbak, N.N. 1974. Dva novih dlya fauni SSSR vida yashcheritsiz semeysteva gekkonov. Vestnik Zoologii (Kiev) 1974 (4): 33-39
  • Khan M S 1991. A new Tenuidactylus gecko from the Sulaiman Range, Punjab, Pakistan. Journal of Herpetology 25 (2): 199-204 - get paper here
  • Khan M S; Tasnim R 1990. A new gecko of the genus Tenuidactylus from northeastern Punjab, Pakistan, and southwestern Azad Kashmir. Herpetologica 46 (2) 1990: 142-148 - get paper here
  • Khan, M.S. 2003. Questions of generic designation of angular-toed geckos of Pakistan with descriptions of three new genera (Reptilia: Gekkonidae). J. nat. hist. Wildl. 2 (2): 1-9
  • Khan, Muhamad Sharif. 2008. Biology and distribution of geckos of genus Indogekko Khan, 2003 (Sauria: Gekkonidae). Russ. J. Herpetol. 15 (2):87-92 - get paper here
  • Lantz, L. A. 1918. Reptiles from the River Tajan (Transcaspia). Proc. Zool. Soc. London 1918: 11—17 - get paper here
  • Leviton A E; Anderson S C 1984. Description of a new species of Cyrtodactylus from Afghanistan with remarks on the status of Gymnodactylus longipes and Cyrtodactylus fedtschenkoi. Journal of Herpetology 18 (3): 270-276 - get paper here
  • Macey,J.R.; Ananjeva, N.B.; Wang, Y. & Papenfuss, T.J. 2000. Phylogenetic relationships among Asian gekkonid lizards formerly of the genus Cyrtodactylus based on cladistic analyses of allozyme data: monophyly of Cyrtopodion and Mediodactylus. Journal of Herpetology 34 (2): 258-265 - get paper here
  • Nikolsky, A. M. 1896. Diagnoses reptilium et amphibiorum novorum in Persia orientali a N. Zarudny collectorum. Annuaire Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg, 1:369-372. - get paper here
  • Nikolsky, A. M. 1897. Les reptiles, amphibiens et poissons recueillis (part.) Mr. N. Zaroudny dans la Perse orientale [in Russian]. Annuaire Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg 2: 306-348 - get paper here
  • Orlov, N. L. 1981. About the eastern range limit of the gecko Gymnodactylus longipes Nikolsky, 1896 (in Russian). Proceedings of the Zoological Institute of the Academy of Sciences, U.S.S.R. 101, 89-91
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Rösler, H. 2000. Die postanale Beschuppung bei Cyrtodactylus GRAY 1827 und Cyrtopodion FITZINGER 1843 - funktionelle und taxonomische Aspekte (Sauria: Gekkonidae). Gekkota 2: 154-207
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, H. 2018. Haftbar -- Die einzigartige Welt der Geckos. Begleitheft zur Ausstellung im Naturkundemuseum Erfurt Mai-Aug 2018, 96 pp.
  • Rösler, Herbert 1995. Geckos der Welt - Alle Gattungen. Urania, Leipzig, 256 pp.
  • Rösler, Herbert 1997. Paläarktische Geckos (Reptilia: Gekkota) Teil I: Historischer Überblick, Geographie und Klima, Verbreitung. Sauria 19 (1): 17-28 - get paper here
  • Shcherbak, N. N. [= Szczerbak], and M. L. Golubev 1996. Gecko fauna of the USSR and contiguous regions. SSAR [Russian original published in 1986]. [book review in Salamandra 36 (2): 138]
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • ŠMÍD, JIŘÍ; JIŘÍ MORAVEC, PETR KODYM, LUKÁŠ KRATOCHVÍL, SEYYED SAEED HOSSEINIAN YOUSEFKHANI, ESKANDAR RASTEGAR-POUYANI & DANIEL FRYNTA 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa 3855 (1): 001–097 - get paper here
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp. - get paper here
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