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Trapelus flavimaculatus RÜPPELL, 1835

IUCN Red List - Trapelus flavimaculatus - Least Concern, LC
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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Yellow-spotted Agama 
SynonymTrapelus flavimaculatus RÜPPELL 1835: 12
Agama flavimaculata — ANDERSON 1896: 31
Agama jayakari ANDERSON 1896: 65 (fide ARNOLD 1980)
Agama flavimaculata — BOULENGER 1919: 112
Agama flavimaculata — LOVERIDGE 1936: 52
Agama jayakari — HAAS 1957: 67
Agama jayakari — WERMUTH 1967: 16
Agama jayakari — LEVITON & ANDERSON 1967: 159
Agama flavimaculata — WERMUTH 1967: 13
Agama flavimaculata — ARNOLD 1986
Trapelus flavimaculatus — JOGER 1987: 259
Trapelus jayakari — ANDERSON 1999
Trapelus jayakari — HENKEL 2003
Trapelus flavimaculatus — BARTS & WILMS 2003
Trapelus jayakari — WAGNER et al. 2011 
DistributionW Saudi Arabia, Oman, United Arab Emirates (UAE), Qatar

Type locality: Djetta, Arabien.

jayakari: Saudi Arabia; Type locality: Maskat  
Reproductionoviparous 
TypesHolotype: BMNH 1920.1.20.1241
Types: BMNH 1946.8.14.86-91 (and possibly additional specimens) []jayakari] 
DiagnosisDiagnosis (jayakari): This species is closely allied to A. flavimaculata, Rüppell, from Jiddah, but is distinguished from it by its large, regular, strongly keeled and mucronate scales, by its less cordate head, which is shorter than the tibia, and by the more strongly keeled character of its caudal scales. It also attains to a greater size, and the scales on the back of the head are somewhat more spinose than in A. flavimaculata, Rüppell. The entire absence of preanal pores and callose scales in these two species necessitates an alteration in the hitherto accepted definition of the genus Agama (Anderson 1896: 67). 
CommentSynonymy: Trapelus jayakari has been synonymized with Trapelus flavimaculatus by Arnold 1986.

Distribution: See map in Burriel-Carranza et al. 2019 for map in UAE. Not in Egypt (El Din 2006). 
EtymologyNamed after Latin “flavus” = yellow and “macula” = spots, for the yellow spots.

Agama jayakari was named after Atmaram S. Jayakar (1844-1911), Indian surgeon, sent to Muscat by the Indian Medical Service. 
References
  • Aloufi, A., Amr, Z., & Baker, M. A. 2022. Reptiles from'Uruq Bani Ma'arid and Harat al Harrah protected areas in Saudi Arabia: Reptiles from two protected areas in Saudi Arabia. Herpetology Notes 15: 483-491
  • Aloufi, Abdulhadi A.; Zuhair S. Amr, Mohammad A. Abu Baker 2021. Reptiles and Amphibians of Al Madinah Al Munawwarah Province, Saudi Arabia. Russian Journal of Herpetology 28 (3): 1 - get paper here
  • Anderson, John 1896. A Contribution to the Herpetology of Arabia, with a preliminary list of the reptiles and batrachians of Egypt. London, R. H. Porter, 124 pp. - get paper here
  • Arnold E N 1980. The scientific results of the Oman flora and fauna survey 1977 (Dhofar). The reptiles and amphibians of Dhofar, southern Arabia. JOURNAL OF OMAN STUDIES SPECIAL REPORT (No. 2): 273-332 - get paper here
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger,G.A. 1919. Sur l’Agama tournevillei Lataste, du Sahara Algérien. Bull. Soc. Zool. France (PARIS) 44: 112 - get paper here
  • Burriel-Carranza B, Pedro Tarroso, Johannes Els, Andrew Gardner, Pritpal Soorae, Ahmed Ali Mohammed, Sai Ravi Krishna Tubati, Mohamed Mustafa Eltayeb, Junid Nazeer Shah, Héctor Tejero-Cicuéndez, Marc Simó-Riudalbas, Juan Manuel Pleguezuelos, Daniel F 2019. An integrative assessment of the diversity, phylogeny, distribution, and conservation of the terrestrial reptiles (Sauropsida, Squamata) of the United Arab Emirates. PLoS ONE 14(5): e0216273 - get paper here
  • Carranza S, Xipell M, Tarroso P, Gardner A, Arnold EN, Robinson MD, et al. 2018. Diversity, distribution and conservation of the terrestrial reptiles of Oman (Sauropsida, Squamata). PLoS One 13 (2): e0190389 - get paper here
  • Carranza, Salvador; Johannes Els; Bernat Burriel-Carranza 2021. A field guide to the reptiles of Oman. Madrid : Consejo Superior de Investigaciones Científicas, 223 pp. [review in HR 53 (3): 531] - get paper here
  • Cogălniceanu, Dan; Aurora Castilla, Aitor Valdeon, Alberto Gosa, Noora Al Jaidah, Ali Alkuwary, Essam Saifelnasr, Paloma Mas, Renee Richer, Ahmad Amer Al Hemaidi 2014. A preliminary report on the distribution of lizards in Qatar. ZooKeys 373 (2014): 67-91<br />doi: 10.3897/zookeys.373.5994 - get paper here
  • Eissa, S.M. & EL ASSY, Y.S. 1975. Record of certain reptilian species found in Kuwait. Jour. Univ. Kuwait (Sci.), 2: 123-146 - get paper here
  • Gallagher, M.D. 1971. Amphibians and Reptiles of Bahrain. pub. privately, Bahrain 40 pp.
  • Gardner, A.S. 2013. The amphibians and reptiles of Oman and the UAE. Edition Chimaira, 480 pp.
  • Haas, Georg 1957. Some amphibians and reptiles from Arabia. Proc. Cal. Acad. Sci. 29 (3): 47-86 - get paper here
  • Henkel, F.-W. 2003. Herpetological expedition through Oman. Reptilia (GB) (27): 50-55 - get paper here
  • Joger,U. 1987. An interpetation of reptile zoogeography in Arabia, with special reference to Arabian herpetofaunal relations with Africa. In: Krupp,F., Schneider,W. & Kinzelbach,R. (eds.): Proceedings of the symposium on the fauna and zoogeography of the Middle Eas Beihefte zum Tübinger Atlas des Vorderen Orients, A 28, Wiesbaden: 257-271.
  • Jongbloed, M. 2000. Field Guide to the reptiles and amphibians of the UAE - Wild about reptiles. Barkers Trident Communications, 116 pp.
  • Leviton,A.E. & Anderson,S.C. 1967. Survey of the reptiles of the Sheikdom of Abu Dhabi, Arabian Peninsula. Part II. Systematic account of the collction of reptiles made in the Sheikdom of Abu Daby by John Gasperetti. Proc. Cal. Acad. Sci. (4) 35: 157-192 - get paper here
  • Leviton,A.E.; Anderson,S.C.; Adler, K.; Minton,S.A. 1992. Handbook to Middle East Amphibians and Reptiles. SSAR, Oxford, Ohio (Contr. to Herpetol. No. 8), 1-252
  • Loveridge, A. 1936. African reptiles and amphibians in the Field Museum of Natural History. Zool. Ser. Field Mus. Nat. Hist., Chicago, 22 (1): 1-122 - get paper here
  • Rüppell, E. 1835. Neue Wirbelthiere zu der Fauna von Abyssinien gehörig, entdeckt und beschrieben. Amphibien. S. Schmerber, Frankfurt a. M. - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, Roberto; Riccardo Nincheri, Benedetto Lanza 2014. Catalogue of Arabian reptiles in the collections of the “La Specola” Museum, Florence. Scripta Herpetologica. Studies on Amphibians and Reptiles in honour of Benedetto Lanza: pp. 137-164 - get paper here
  • van der Kooij, Jeroen 2001. The herpetofauna of the Sultanate of Oman: Part 1: The amphibians, worm lizards, agamas and chameleons. Podarcis 1 (3): 71-83 - get paper here
  • WAGNER, PHILIPP; JANE MELVILLE, THOMAS M. WILMS and ANDREAS SCHMITZ 2011. Opening a box of cryptic taxa – the first review of the North African desert lizards in the Trapelus mutabilis Merrem, 1820 complex (Squamata: Agamidae) with descriptions of new taxa. Zoological Journal of the Linnean Society 163: 884–912 - get paper here
  • Wilms, T. 2007. Unternehmen “Dornschwanzagame”: Das Reptilium-Freilandforschungsprojekt in Saudi-Arabien. Draco 8 (31): 45-54 - get paper here
 
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