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Trapelus savignii (DUMÉRIL & BIBRON, 1837)

IUCN Red List - Trapelus savignii - Vulnerable, VU
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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common Names 
SynonymAgama savignii DUMÉRIL & BIBRON 1837: 508
Phrynopsis savignyi FITZINGER 1843: 80 (nom. subst.)
Agama savignyi — BARASH & HOOFIEN 1961: 103
Agama savignii — WERMUTH 1967: 23
Trapelus savignii — MACEY et al. 2000
Trapelus savignyi — MÜLLER 2001
Trapelus savignii — BARTS & WILMS 2003
Trapelus savignyi — SINDACO & JEREMČENKO 2008
Trapelus savignii — MEIRI et al. 2019 
DistributionEgypt (incl. Sinai), Israel

Type locality: Egypte; restricted to El Arish, northern Sinai, Egypt, by neotype designation.  
Reproductionoviparous 
TypesNeotype: ZFMK 77470, designated by WAGNER & CROCHET 2009. 
DiagnosisDiagnosis: Diagnosis: Medium-sized Trapelus [largest known voucher from Egypt SVL 123 mm (Anderson, 1898)] with an average ratio TL/SVL of 1.3. Gular pouch well developed. Dorsal scales relatively uniform with intermixed larger scales. Ventral scales keeled. Usually vertebral scales slightly larger than the scales on the flanks. Coloration uniformly sandy grey with five brown transverse bands, which are inter- rupted by a yellowish stripe and whitish lines on the body and a barred tail. Additionally two transverse bands are on the interorbital area. Adult males get a characteristic coloration under nuptial conditions: throat and flanks bright to violet blue, the flanks with interspersed white spots. Additionally, pregnant females have a special coloration, with dorsal bands becoming brick red (from Wagner et al. 2011: 904). 
CommentSynonymy after WERMUTH 1967.

Distribution: see map in Kissling et al. 2016: Fig. 1. Not in Israel where it is replaced by P. agnetae. 
References
  • Baha El Din, Sherif M. and Omar Attum 2000. The herpetofauna of Zaranik Protected Area, Egypt, with notes on their ecology and conservation. Herpetological Bulletin (73): 17-21 - get paper here
  • Bar, Aviad and Guy Haimovitch 2012. A Field Guide to Reptiles and Amphibians of Israel. Pazbar LTD, 246 pp. - get paper here
  • Barash, Al & Hoofien, J.H. 1956. Reptiles of Israel. [in Hebrew] Tel Aviv, 179 pp. [also cited as 1961]
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Fitzinger, L. 1843. Systema Reptilium, fasciculus primus, Amblyglossae. Braumüller et Seidel, Wien: 106 pp. - get paper here
  • Goldberg, Stephen R. 2013. Trapelus savignii (Egyptian sand agama) reproduction. Herpetological Review 44 (4): 685-686 - get paper here
  • Gorman, George C.;Shochat, Dan 1972. A taxonomic interpretation of chromosomal and electrophoretic data on the agamid lizards of Israel with notes on some East African species. Herpetologica 28 (2): 106-112 - get paper here
  • Ibrahim, Adel A. 2013. The Herpetology of the Suez Canal Zone, Egypt. Vertebrate Zoology 63 (1): 87–110 - get paper here
  • Kissling, W. Daniel; Anne Blach-Overgaard, Roelof E. Zwaan & Philipp Wagner 2016. Historical colonization and dispersal limitation supplement climate and topography in shaping species richness of African lizards (Reptilia: Agaminae). Scientific Reports 6: 34014, doi:10.1038/srep34014 - get paper here
  • Macey, J. R., J. A. Schulte II, A. Larson, N. B. Ananjeva, Y. Wang, R. Pethiyagoda, N. Rastegar-Pouyani, T. J. Papenfuss 2000. Evaluating trans-Tethys migration: an example using acrodont lizard phylogenetics. Systematic Biology 49 (2): 233-256 - get paper here
  • Marx,Hymen 1968. Checklist of the reptiles and amphibians of Egypt. Spec. Publ. U.S. Nav. Med. Res. Unit 3: 1-91
  • Meiri, Shai; Amos Belmaker, Daniel Berkowic, Kesem Kazes, Erez Maza, Guy Bar-Oz and Roi Dor 2019. A checklist of Israeli land vertebrates. Israel Journal of Ecology and Evolution - get paper here
  • Müller, H. D. 2001. Trapelus savignyi (DUMÉRIL & BIBRON 1837) - Erfahrung mit der Terrarienhaltung. Elaphe 9 (4): 7-16
  • NÚÑEZ, Herman; DANIEL PINCHEIRA-DONOSO 2006. Liolaemus confusus, a new species of lizard for the coastal range of central Chile (Sauria, Liolaeminae): phenotypic and cytogenetic evidence. Bol Mus Nac Hist Nat Bol Chile 55-56: 77
  • Schleich, H.H., Kästle,W., Kabisch, K. 1996. Amphibians and Reptiles of North Africa. Koeltz, Koenigstein, 627 pp.
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Wagner, P. & Böhme, W. 2006. A new species of the genus Trapelus Cuvier, 1816 (Squamata: Agamidae) from arid central Africa. Bonner Zoologische Beiträge 55 (2): 81-87 - get paper here
  • Wagner, P. & Crochet, P.A. 2009. The status of the nomina Trapelus savignyi Audouin, 1827 and Agama savignii Duméril & Bibron, 1837 and the valid nomen of the Savigny’s Agama (Sauria: Agamidae). Zootaxa 2209: 57–64 - get paper here
  • WAGNER, PHILIPP; JANE MELVILLE, THOMAS M. WILMS and ANDREAS SCHMITZ 2011. Opening a box of cryptic taxa – the first review of the North African desert lizards in the Trapelus mutabilis Merrem, 1820 complex (Squamata: Agamidae) with descriptions of new taxa. Zoological Journal of the Linnean Society 163: 884–912 - get paper here
 
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