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Tropidonophis doriae (BOULENGER, 1898)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Barred keelback 
SynonymTropidonotus doriae BOULENGER 1898: 704
Tropidonotus doriae — LIDTH DE JEUDE 1911: 272
Natrix doriae — LOVERIDGE 1948: 381
Amphiesma doriae — MALNATE 1960: 50
Tropidonophis doriae — MALNATE & UNDERWOOD 1988: 102
Tropidonophis doriae — READ 1998
Tropidonophis (Desburkeus) doriae desburkei HOSER 2012
Tropidonophis doriae — WALLACH et al. 2014: 749 
DistributionIndonesia (Aru, Irian Jaya),
Papua New Guinea

Type locality: Moroka, 4200 feet, Bartholomew Range, Central Province, Papua New Guinea [BMNH 97.12.10.116]; Haveri, 2300 feet, Bartholomew Range, Centra Province, Papua New Guinea [BMNH 97.12.10.117-118]  
Reproductionoviparous 
TypesSyntypes: BMNH 97.12.10.116; BMNH 97.12.10.117-118; MRSN (MSNTO) R946, MSNG 36101 (3 specimens). Remaining 11 syntypes (from Haveri and Moroka) not traced. 
DiagnosisDescription (based on 18 specimens): Dentition as in T. picturatus and T. mairii (25-29 maxillary teeth). Eye rather large. Rostral twice as broad as deep, just visible from above; internasals as long as broad or a little longer than broad, broadly truncate in front , as long as the praefrontals; frontal once and a half to once and two thirds as long as Inroad, longer than its distance from the end of the snout, shorter than the parietals; loreal deeper than long; two praeand three postoculars; temporals usually 2 + 2 sometimes 1 + 2 or 1 + 1; eight (exceptionally nine) upper labials, third, fourth and fifth (or fourth, fifth and sixth) entering the eye; four or five lower labials in contact with the anterior chin-shields, which are shorter than the posteriors. Scales in 17 rows, strongly keeled, outer rou feebly keeled. Ventrals 146-158; anal divided; subcaudals 69-79 (Boulenger 1898: 704).

Coloration: Yellowish, reddish, or dark grey above, with more or less distinct black or blackish cross-bars as in T. mairii, var. semicinctus, D. & B.; upper labials yellowish, with black sutures; yellowish beneath, uniform or with some darker dots, none of the shields with dark edges (Boulenger 1898: 704).

Size: Total length 990 mm.; tail 250 (Boulenger 1898: 704).

Detailed Description: Malnate & Underwood 1988: 102. 
CommentSynonymy after MALNATE & UNDERWOOD 1988 
EtymologyNamed after Marchese Giacomo Doria (1840-1913), Italian zoologist who collected in Persia (Iran) with de Filippi (1862-1863) and in Borneo with Beccari (1865-1866). He founded the Museo Civico di Storia Naturale, Turin (1867-1913), and was its first Director. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G. A. 1898. An account of the reptiles and batrachians collected by Dr. L. Loria in British New Guinea. Annali del Museo Civico di Storia Naturale di Genova, 18: 694—710 [1897?] - get paper here
  • Clegg, Jonathan R. and Merlijn Jocque 2015. The Collection of Snakes Made by Benoît Mys and Jan Swerts in Northern Papua New Guinea in 1982–85. Journal of Herpetology 50 (3): 476-485 [2016] - get paper here
  • Goldberg, Stephen R. and Charles R. Bursey. 2011. Tropidonophis doriae (barred keelback) endoparasites. Herpetological Review 42 (3): 447 - get paper here
  • Hoser, R.T. 2012. A review of natricine genera Tropidonophis Jan, 1863 and Amphiesma Duméril, Bibron and Duméril, 1854 (Serpentes: Colubroidae [sic]: Natricinae). Australasian J. Herpetol. 13:35–46. - get paper here
  • Kaiser, H.; Crother, B.I.; Kelly, C.M.R.; Luiselli, L.; O’Shea, M.; Ota, H.; Passos, P.; Schleip, W.D. & Wüster, W. 2013. Best Practices: In the 21st Century, Taxonomic Decisions in Herpetology are Acceptable Only When Supported by a Body of Evidence and Published via Peer-Review. Herpetological Review 44 (1): 8-23
  • Lidth DE JEUDE, T.W. VAN 1911. Reptilien (Schlangen). Nova Guinea. Résultats de l’expedition scientifique néerlandaise à la Nouvelle Guinée en 1907 sous les auspices de Dr. H.A. Lorenz 9. Leiden (E. J. Brill), pp. 265-287
  • Loveridge, A. 1948. New Guinean reptiles and amphibians in the Museum of Comparative Zoology and United States National Museum. Bull. Mus. Comp. Zool. Harvard 101 (2): 305-430. - get paper here
  • Malnate E V; Underwood G 1988. Australasian natricine snakes of the genus Tropidonophis. PROCEEDINGS OF THE ACADEMY OF NATURAL SCIENCES OF PHILADELPHIA 140 (1) : 59-201 - get paper here
  • Malnate, Edmond V. 1960. Systematic division and evolution of the colubrid snake genus Natrix, with comments on the subfamily Natricinae. Proc. Acad. Nat. Sci. Philadelphia, 112 (3): 41-71. - get paper here
  • O'Shea,M. 1996. A Guide to the Snakes of Papua New Guinea. Independent Publishing, Port Moresby, xii + 239 pp. - get paper here
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Read, John L. 1998. Reptiles and amphibians of the Kau Wildlife Area near Madang: a valuable conservation resource. Science in New Guinea 23 (3): 145-152
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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