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Tukutuku rakiurae (THOMAS, 1981)

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Higher TaxaDiplodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Harlequin gecko 
SynonymHoplodactylus rakiurae THOMAS 1981
Hoplodactylus rakiurae — KLUGE 1993
Hoplodactylus rakiurae — RÖSLER 1995: 71
Tukutuku rakiurae — NIELSEN et al. 2011
Tukutuku kurakiurae — BAUER 2013 (plates; in error) 
DistributionNew Zealand (Southern Stewart Island)

Type locality: southern Tin Range, Stewart Island (47°8'30"S, 167°44'30"E).  
TypesHolotype: NMNZ R1825, adult female; paratypes: NMNZ 
DiagnosisDefinition (genus). Pupils with crenulated margins; scansorial pads of digits II–V not continuous along length of entire digit, a short series of fragmented scales separate lamellae from claw base; toes narrow, with straight lamellae; apical plates present at the base of claws of all digits of the pes; ventral skin opaque; dorsal body pattern with complex chevron pattern; mouth lining and tongue pigmented; precloacal pores of males continuous with femoral pores on under surface of thighs; SVL to 71 mm [NIELSEN et al. 2011]. 
CommentDistribution: Gecko with one of the most southern distribution (except some South American species which may get further south).

Type species: Tukutuku rakiurae (Thomas, 1981) is the type species of the genus Tukutuku NIELSEN et al. 2011. 
EtymologyEtymology (genus): A Maori word for ornamental lattice-work, often intricate and beautiful, referring to the sole species’ unique dorsal coloration pattern. Maori nouns typically have no gender, but we treat Tukutuku as masculine for the purposes of nomenclatural construction. The specific name is derived from “Rakiura”, the Maori name for Stewart Island. 
  • Bauer, A.M. 2013. Geckos - The Animal Answer Guide. Johns Hopkins University Press, 159 pp.
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Henle, K. 1981. Die Herpetofauna Neuseelands, Teil 4: Endemische Geckos. Herpetofauna 3 (15): 16-19 - get paper here
  • Hitchmough, Rodney A.; Geoffrey B. Patterson, and David G. Chapple 2016. Putting a Name to Diversity: Taxonomy of the New Zealand Lizard Fauna. in: Chapple, D.G. (ed). New Zealand Lizards. Springer, pp. 87-108 - get paper here
  • Jewell, Tony 2008. Hoplodactylus rakiurae: New Zealand's Enigmatic Harlequin. Gekko 5 (2): 19-24
  • Macredie, Grant 1998. The Harlequin gecko Hoplodactylus rakiurae (Thomas 1980). Dactylus 3 (2): 61-62
  • Nielsen, Stuart V.; Aaron M. Bauer, Todd R. Jackman, Rod A. Hitchmough and Charles H. Daugherty 2011. New Zealand geckos (Diplodactylidae): Cryptic diversity in a post-Gondwanan lineage with trans-Tasman affinities. Molecular Phylogenetics and Evolution 59 (1): 1-22 - get paper here
  • Rösler, Herbert 1995. Geckos der Welt - Alle Gattungen. Urania, Leipzig, 256 pp.
  • Rösler, Herbert 2017. Gecko-Chorologie (Squamata: Gekkota). Gekkota (4): 1-160
  • Thomas, B.W. 1981. Hoplodactylus rakiurae n. sp. (Reptilia: Gekkonidae) from Stewart Island, New Zealand and comments on the taxonomic status of Heteropholis nebulosus McCann. New Zealand Journal of Zoology 8: 33- 47 - get paper here
  • van Winkel, D., Baling, M. & Hitchmough, R. 2018. Reptiles and Amphibians of New Zealand: A field guide. Auckland University Press, Auckland, 376 pp
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