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Anolis marsupialis TAYLOR, 1956

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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymAnolis humilis marsupialis TAYLOR 1956: 95
Anolis humilis marsupialis — SAVAGE 2002
Anolis marsupialis — SAVAGE & BOLANOS 2009
Anolis marsupialis — KÖHLER et al. 2015
Norops marsupialis — NICHOLSON et al. 2018 
DistributionCosta Rica

Type locality: “about 15 km WSW of San Isidro del General along Dominical Road” (probably Puntarenas Province or near the border of Puntarenas and San José Province).  
Reproductionoviparous 
TypesHolotype: KU (= KUMNH); other specimens: SMF, LACM, MCZ, AUM, UCR, 
DiagnosisDiagnosis: Similar to humilis humilis but larger (to 44 mm.); the enlarged dorsal scales in ten to fourteen rows, the two median rows elevated into a suggestion of a dorsal crest and continuous with a similar nuchal crest, and larger than adjoining scalerows; dorsal scales about equal to or a little larger than ventral scales; axillary pocket much enlarged, the opening nearly as great as its depth, the side walls usually partly scaled; ventrals keeled. (Taylor 1956)


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CommentSpecies group: Norops auratus Species Group (fide Nicholson et al. 2012)

Distribution: For a map of localities see Phillips et al. 2015: Fig. 2, Köhler et al. 2025: 119 (Fig. 8). 
EtymologyNamed after Latin marsupium, or Greek μαρσίπιον, pouch, purse + Latin -alis, adjectival particle. [“...From Greek word marsipion or Latin, marsupium = pouch bag, in reference to the axillary pouches...”]. 
References
  • KÖHLER, JOHANNES J.; STEVEN POE, MASON J. RYAN & GUNTHER KÖHLER 2015. Anolis marsupialis Taylor 1956, a valid species from southern Pacific Costa Rica (Reptilia, Squamata, Dactyloidae). Zootaxa 3915 (1): 111–122 - get paper here
  • Nicholson KE, C. Guyer, and JG Phillips 2017. Biogeographic Origin of Mainland Norops (Squamata: Dactyloidae). Assumptions Inhibiting Progress in Comparative Biology (eds. Crother and Parenti), pp. 169–184
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Phillips, JG, J. Deitloff, C. Guyer, S. Huetteman, and KE Nicholson 2015. Biogeography and evolution of a widespread Central American lizard species complex: Norops humilis, (Squamata: Dactyloidae). BMC Evolutionary Biology 15: 143 - get paper here
  • Poe, Steven 2025. A Guide to the Anolis Lizards (Anoles) of Mainland Central and South America. Princeton University Press, 432 pp. - get paper here
  • Sasa, M., Arias, E., & Chaves, G. 2025. Annotated list of amphibians and reptiles of Costa Rica: The role of the Museum of Zoology in cataloging the country’s herpetological diversity. Revista de Biología Tropical, 73(S2): e64536-e64536 - get paper here
  • Savage, J.M. 2002. The Amphibians and Reptiles of Costa Rica: A Herpetofauna Between Two Continents, Between Two Seas. University of Chicago Press, 934 pp. [review in Copeia 2003 (1): 205]
  • Savage, J.M. & Bolaños, F. 2009. A checklist of the Amphibians and Reptiles of Costa Rica: Additions and nomenclatural revisions. Zootaxa 2005: 1–23 - get paper here
  • Taylor, E. H. 1956. A review of the lizards of Costa Rica. Univ. Kansas Sci. Bull. 38 (part 1): 3-322 - get paper here
 
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