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Enyalius pictus (SCHINZ, 1822)

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Higher TaxaLeiosauridae (Enyaliinae), Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesPortuguese: Camaleãozinho, Papa-Vento 
SynonymAgama picta SCHINZ 1822
Agama picta — WIED-NEUWIED 1825
Ophryessa margariticeus — BOIE 1825
Ecphymotes pictus — FITZINGER 1826: 49
Uraniscodon (Pneustes) picta — KAUP 1827: 612
Hypsibatus picta — WAGLER 1830: 150
Hypselophus picta — WIEGMANN 1835
Ophyessa (Xiphura) margariticeus — GRAY 1831: 40
Calotes pictus — SCHINZ 1835: 86
Uperanodon pictum — DUMÉRIL & BIBRON 1837: 251
Hypsibatus (Uperanodon) pictus — FITZINGER 1843
Uperanodon Pictum — DUMÉRIL, BIBRON & DUMÉRIL 1854: 268
Enyalius zonatus WETTSTEIN 1926: 1 (see comment)
Enyalius pictus — ETHERIDGE 1969: 240
Enyalius pictus — PETERS & DONOSO-BARROS 1970: 118
Enyalius catenatus pictus — JACKSON 1978
Enyalius pictus — FROST et al. 2001
Enyalius catenatus pictus — RODRIGUES et al. 2006
Enyalius pictus — RODRIGUES et al. 2014
Enyalius pictus — CARVALHO et al. 2023 
DistributionBrazil (Bahia, Espírito Santo)

Type locality: Mucurí and Lago d’Arara, Bahia, Brazil.  
Reproductionoviparous 
TypesNeotype: NMW 13894, from “Muriti dist.”, designated by ETHERIDGE 1968. Holotype was AMNH 13894, but now lost (ETHERIDGE 1968) 
DiagnosisDiagnosis: This subspecies is distinguished from others of the species by the following characters: 1) enlarged supraoculars not keeled; 2) enlarged supraoculars as large as or larger than scales of the supraorbital series; 3) ventral scales smooth; 4) infratibials smooth or with weak keels; 5) infratibials wider than long; 6) one of the suboculars enlarged or not; 7) keel of preocular medially located; 8) midbody scale count greater than 140; 9) shank/snout-vent length ratio greater than .30. (JACKSON 1978: 21)


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CommentSynonymy: JACKSON (1978) suggested that E. bibronii, E. pictus and E. catenatus were intergrading subspecies of on species. Synonymy partly after PETERS & DONOSO-BARROS 1970, ETHERIDGE 1969. The author of this species is SCHINZ 1822, not WIED 1823 (or 1825) fide MYERS et al. 2011.

References: The paper of WETTSTEIN (1926) is cited in PETERS & DONOSO-BARROS 1970 as being published in “Anz. Acad. Wiss. Wien 63: 1”.

The original color plate of WIED-NEUWIED (1825) is reproduced in RODRIGUES et al. (2006).

Distribution: Not in Ecuador fide TORRES-CARVAJAL 2001, TORRES-CARVAJAL et al. 2019. See map in Breitman et al. 2019: 357 (Fig. 1). 
EtymologyNamed after Latin pictus, painted; colored; decorated. (SCHINZ 1822, Esteban Lavilla, pers. comm., April 2024) 
References
  • Breitman, M. Florencia; Fabricius M.C.B. Domingos, Justin C. Bagley, Helga C. Wiederhecker, Tayná B. Ferrari, Vitor H.G.L. Cavalcante, André C. Pereira, Tarcísio L.S. Abreu, Anderson Kennedy Soares De-Lima, Carlos J.S. Morais, Ana C.H. del Prette, Iz 2018. A New Species of Enyalius (Squamata, Leiosauridae) Endemic to the Brazilian Cerrado. Herpetologica 74 (4): 355-369. - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Duméril, A. M.C., G. BIBRON & A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 9. Paris, XX + 440 S. - get paper here
  • Duméril, A.M.C., and G. Bibron. 1835. Erpétologie Générale ou Histoire Naturelle Complète des Reptiles, Vol. 2. Librairie Encyclopédique de Roret, Paris, iv + 680 p. - get paper here
  • Etheridge, R. 1969. A review of the Iguanid lizard genus Enyalius. Bulletin of the British Museum (Natural History), Zoology 18 (8): 233-260 - get paper here
  • Frost, Darrel R.; Richard Etheridge; Daniel Janies and Tom A. Titus 2001. Total evidence, sequence alignment, evolution of polychrotid lizards, and a reclassification of the Iguania (Squamata, Iguania). American Museum Novitates 3343: 1-22 - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248 - get paper here
  • Gonzalez R. C. et al. 2020. Lista dos Nomes Populares dos Répteis no Brasil – Primeira Versão. Herpetologia Brasileira 9 (2): 121 – 214 - get paper here
  • Jackson J F 1978. Differentiation in the genera Enyalius and Strobilurus (Iguanidae): implications for Pleistocene climatic changes in eastern Brazil. Arquivos de Zoologia 30(1) 1978: 1-79
  • Peters, James A. & Donoso-Barros, Roberto 1970. Catalogue of the Neotropical Squamata: Part II. Lizards and Amphisbaenians. Bull. US Natl. Mus. 297: 293 pp. - get paper here
  • Rodrigues, Miguel Trefaut; Carolina Elena Viña Bertolotto, Renata Cecília Amaro, Yatiyo Yonenaga-Yassuda, Eliza Maria Xavier Freire, Katia Cristina Machado Pellegrino 2014. Molecular phylogeny, species limits, and biogeography of the Brazilian endemic lizard genus Enyalius (Squamata: Leiosauridae): An example of the historical relationship between Atlantic Forests and Amazonia. Molecular Phylogenetics and Evolution 81: 137-146, DOI: 10.1016/j.ympev.2014.07.019 - get paper here
  • Rodrigues, Miguel Trefaut; Marco Antonio de Freitas, Thais Figueiredo Santos Silva, and Carolina Elena Viña Bertolotto 2006. A new species of lizard genus Enyalius (Squamata, Leiosauridae) from the highlands of Chapada Diamantina, State of Bahia, Brazil, with a key to species. Phyllomedusa 5 (1): 11-24 - get paper here
  • Schinz, H. R. 1822. Das Thierreich eingetheilt nach dem Bau der Thiere als Grundlage ihrer Naturgeschichte und der vergleichenden Anatomie von dem Herrn Ritter von Cuvier. Vol. 2. J. G. Cotta, Stuttgart. - get paper here
  • Uchôa LR, Delfim FR, Mesquita DO, Colli GR, Garda AA, Guedes TB 2022. Lizards (Reptilia: Squamata) from the Caatinga, northeastern Brazil: Detailed and updated overview. Vertebrate Zoology 72: 599-659 - get paper here
  • Wettstein, Otto von 1926. Eine neue Eidechse der Gattung Enyalis aus Ecuador. Akademischen Anzeiger (1): 1-2
  • Wied-Neuwied, M. Prinz zu 1824. Abbildungen zur Naturgeschichte von Brasilien. [1822-1831] Weimar - get paper here
 
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