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Tropidonophis dahlii (WERNER, 1899)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: New Britain keelback 
SynonymTropidonotus (Macropophis) dahlii WERNER 1899: 373
Tropidonotus hypomelas STERNFELD 1913: 386 (part.)
Tropidonophis (Macropophis) melanocephalus WERNER 1925: 47
Natrix melanocephala — LOVERIDGE 1948: 380
Macropophis hypomela — MALNATE 1960: 47 (part.)
Macropophis melanocephala — MALNATE 1960: 47 (part.)
Tropidonophis dahlii — MALNATE & UNDERWOOD 1988: 169
Tropidonophis dahlii — WALLACH et al. 2014: 749
Tropidonophis dahlii — ROBERTS et al. 2024 
DistributionIndonesia (Irian Jaya, Otakwa River), Papua New Guinea: Bismarck Archipelago

Type locality: Herbertsgöhe, Neu-Pommern [= New Britain]  
Reproductionoviparous 
TypesLectotype: NMW = NHMV 23488, female (designated by MALNATE & UNDERWOOD 1988) 
DiagnosisDiagnosis: Tropidonophis dahlii is distinguished from hypomelas by the following characters: (1) fewer ventrals (170-185 vs 184-196 in hypomelas) and, apparently, more subcaudals (101-114 vs 103-104 in two hypomelas with complete tails); the v+sc appears similar in the two species but the ratio of subcaudals in the sum is notably higher in dahlii (Table 2); (2) the trachea cartilages continue through the vascular portion of the right lung, extending beyond the anterior end of the liver; in hypomelas the cartilages do not continue beyond the tip of the heart. In addition, the data suggest that the hemipenes of dahlii are somewhat longer and have a deeper fork than the organs of hypomelas (Table 5). (Malnate & Underwood 1988: 169)


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CommentSynonymy: partly after Malnate & Underwood 1988. A specimen in the Vienna Museum (NHMV 23488), presumably Werner's "cotype", is catalogued there as the holotype of dahlii (Tiedemann and Häupl 1980). 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Clegg, Jonathan R. and Merlijn Jocque 2015. The Collection of Snakes Made by Benoît Mys and Jan Swerts in Northern Papua New Guinea in 1982–85. Journal of Herpetology 50 (3): 476-485 [2016] - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248 - get paper here
  • Loveridge, A. 1948. New Guinean reptiles and amphibians in the Museum of Comparative Zoology and United States National Museum. Bull. Mus. Comp. Zool. Harvard 101 (2): 305-430. - get paper here
  • Malnate E V; Underwood G 1988. Australasian natricine snakes of the genus Tropidonophis. PROCEEDINGS OF THE ACADEMY OF NATURAL SCIENCES OF PHILADELPHIA 140 (1) : 59-201 - get paper here
  • Malnate, Edmond V. 1960. Systematic division and evolution of the colubrid snake genus Natrix, with comments on the subfamily Natricinae. Proc. Acad. Nat. Sci. Philadelphia, 112 (3): 41-71. - get paper here
  • O’Shea,M. 1996. A Guide to the Snakes of Papua New Guinea. Independent Publishing, Port Moresby, xii + 239 pp. - get paper here
  • Roberts, J. R., Kraus, F., Allison, A., Richards, S. J., Iova, B., Tjaturadi, B., ... & Austin, C. C. 2024. Systematics and biogeo graphy of a Sunda-Papuan snake lineage (Natricidae: Tropidonophis Jan 1863). Zoological Journal of the Linnean Society, zlae039 - get paper here
  • Sternfeld, R. 1913. Beiträge zur Schlangenfauna Neuguineas und der benachbarten Inselgruppen. Sber. Ges. naturf. Berlin 1913: 384-388
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Werner, F. 1899. Beiträge zur Herpetologie der pacifischen Inselwelt und von Kleinasien. I. Bemerkungen über einige Reptilien aus Neu-Guinea und Polynesien. II. Über einige Reptilien und Batrachier aus Kleinasien. Zool. Anz. 22: 371-375, 375-378 - get paper here
  • Werner. F. 1925. Neue oder wenig bekannte Schlangen aus dem Naturhistorischen Staatsmuseum in Wien. II. Teil. Sitzungsb. Akad. Wiss., Wien, Abt. l, 134: 45 - 66 - get paper here
 
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